Proterotherioidea
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Proterotherioidea
Proterotheriidae is an extinction, extinct family of fossil ungulates from the Cenozoic era that displays toe reduction. Despite resembling primitive, small horses, they were only distantly related to them, and instead belonged to the native South American ungulate order Litopterna. Description Two subfamilies and 18 genera of Proterotheriidae are known. All forms were small or medium-sized. Typical is a reduction of the number of toes and brachydont or mesodont teeth. The family is recorded since the late Palaeocene. Various fossils are known from many parts of the South American continent. The diversity decreased in the Miocene to Pliocene and it has been assumed for a long time that they entirely disappeared in the late Pliocene. However, fossils found in Argentina, Brazil and Uruguay show that one member of the group, ''Neolicaphrium recens'' survived into the Late Lujanian, Pleistocene. Better known genera of the family include ''Diadiaphorus'' and ''Thoatherium'' from th ...
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Litopterna
Litopterna (from grc, λῑτή πτέρνα "smooth heel") is an extinct order of fossil hoofed mammals from the Cenozoic era. The order is one of the five great orders of South American ungulates that were endemic to the continent, until the Great American Biotic Interchange brought new ungulate species. Like other endemic South American mammals, their relationship to other mammal groups had long been unclear, but recent genetic and proteomic evidence indicates that their closest living relatives are Perissodactyls (odd-toed ungulates) including horses, rhinoceros, and tapirs, and that litopterns are closely related to notoungulates, another widespread group of South American ungulates. There were two major groups of litopterns: Proterotheriidae and Macraucheniidae. Proterotheriids were medium to large animals that evolved adaptations for fast running, and occupied a variety of niches that elsewhere were filled by animals such as goats and antelopes, mouse deer, and horses. ...
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Late Paleocene
The Thanetian is, in the International Commission on Stratigraphy, ICS Geologic timescale, the latest age (geology), age or uppermost stage (stratigraphy), stratigraphic stage of the Paleocene epoch (geology), Epoch or series (stratigraphy), Series. It spans the time between . The Thanetian is preceded by the Selandian Age and followed by the Ypresian Age (part of the Eocene). The Thanetian is sometimes referred to as the Late Paleocene. Stratigraphic definition The Thanetian was established by Switzerland, Swiss geologist Eugène Renevier in 1873. The Thanetian is named after the Thanet Formation, the oldest Cenozoic deposit of the London Basin, which was first identified in the area of Kent (southern England) known as the Isle of Thanet. The base of the Thanetian Stage is laid at the base of magnetic chronozone C26n. The references profile (Global Boundary Stratotype Section and Point) is in the Zumaia section (43° 18'N, 2° 16'W) at the beach of Itzurun, Pais Vasco, northern ...
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Brachytherium
''Brachytherium'' is an extinct genus of proterotheriid mammal from the Late Miocene to Late Pliocene of Argentina. It is represented by the type and only species ''B. cuspidatum'', a taxon named in 1883 by Ameghino for a partial mandible with teeth. Though it was considered a dubious taxon at times, ''Brachytherium'' was revised as valid by Schmidt in 2015, who also synonymized the species ''Proterotherium gradatum'' and ''Lophogonodon paranensis'', expanding the material known, all of which is from the Ituzaingó Formation The Ituzaingó Formation ( es, Formación Ituzaingó), in older literature also described as Entre Ríos or Entrerriana Formation, is an extensive geological formation of Late Miocene (Tortonian, or Huayquerian in the SALMA classification) age in .... Some material previously referred to ''Brachytherium'' has been given the new name '' Neobrachytherium''. References {{Taxonbar, from=Q111828098 Proterotheriids Miocene mammals of South America Pliocen ...
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Olisanophus
''Olisanophus'' is a genus of extinct litoptern from the late middle Miocene of southern Bolivia. It was named in 2020 by Andrew McGrath and colleagues, for two distinct species from the same deposits of an unnamed formation of the Honda Group. The type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ... is ''O. riorosarioensis'', known from elements of the partial upper and lower left and right dentitions and possibly a partial mandible, and the referred species is ''O. akilachuta'', known for 6 teeth. Some intermediate material from the same deposits was referred to ''Olisanophus'' sp., not showing diagnostic features of either species. Both species were recovered together in a phylogenetic analysis, where they were sister taxa to a group of '' Diplasiotherium'' and '' Mes ...
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Neodolodus
''Neodolodus'' is a genus of extinct litoptern from the late middle Miocene of southern Colombia. It was named in 1986 by Hoffstetter and Soria, for the species ''Neodolodus colombianus'' from the Castilletes Formation and the La Victoria and Villavieja Formations of the Honda Group. The type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ... ''N. colombianus'' is known from a right mandibular ramus, teeth, and part of the fore- and hindlimbs. ''Neodolodus'' has been classified as a species of '' Prothoatherium'' or '' Lambdaconus'', but was recognized as a distinct genus by McGrath and colleagues in 2020. The taxon was recovered in a phylogenetic analysis as the sister taxon of '' Protheosodon'', between other proterotheriid genera like '' Picturotherium'' and '' Anisolop ...
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Neobrachytherium
''Neobrachytherium'' is an extinct genus of proterotheriid mammal from the Late Miocene of Argentina and Uruguay. It is represented by multiple species, including the type ''N. intermedium'', originally named in 1891 by Moreno and Mercerat as a species of '' Licaphrium'', ''N. morenoi'', originally named in 1914 by Rovereto as a species of ''Brachytherium'', and ''N. ameghinoi'' and ''N. ullumense'', named in 2001 by Soria, who reclassified all the species in the new genus ''Neobrachytherium''. The various species are known from cranial and dental material from the Corral Quemado, Loma de las Tapias and Ituzaingó Formations. ''Neobrachytherium'' may be closely related to ''Thoatherium ''Thoatherium'' (meaning "active swift-beast") is an extinct genus of litoptern mammals from the Early Miocene of Argentina. Fossils of the genus have been found in the Santa Cruz Formation in Argentina.
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Mesolicaphrium
''Mesolicaphrium'' is a genus of extinct litoptern from the late middle Miocene of southern Colombia. It was named in 2020 by Andrew McGrath and colleagues, for the species previously classified as '' Prolicaphrium sanalfonensis'' from the La Victoria and Villavieja Formations of the Honda Group, Colombia. The type species is ''M. sanalfonense'', known from the jaw symphysis and two right mandibular rami, and teeth. ''Mesolicaphrium'' is derived from the similarity to the names ''Prolicaphrium'' and ''Neolicaphrium'', and being in between both taxa in age. The taxon was recovered in a phylogenetic analysis as the sister taxon of '' Diplasiotherium'', closely related to ''Olisanophus ''Olisanophus'' is a genus of extinct litoptern from the late middle Miocene of southern Bolivia. It was named in 2020 by Andrew McGrath and colleagues, for two distinct species from the same deposits of an unnamed formation of the Honda Group. ...''. References {{Meridiungulata, L. Litopterns ...
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Lambdaconus
''Lambdaconus'' is a genus of proterotheriid from the Late Oligocene to Early Miocene of Argentina. The type species is ''L. suinus'', named in 1897 by Ameghino, with referred species including ''L. lacerum'', named as ''Proterotherium ''Proterotherium'' (meaning "first beast") is an extinct genus of litoptern mammal of the family Proterotheriidae that lived during the Late Miocene of Argentina and Chile. Fossils of this genus have been found in the Ituzaingó Formation of Arge ... lacerum'' in 1902 by Ameghino, and ''L. inaqeuifacies''. References {{Taxonbar, from=Q111838995 Proterotheriids Oligocene mammals of South America Miocene mammals of South America Paleogene Argentina Neogene Argentina Fossils of Argentina Colhuehuapian Deseadan Fossil taxa described in 1897 Taxa named by Florentino Ameghino Prehistoric placental genera Golfo San Jorge Basin Sarmiento Formation ...
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Guilielmofloweria
''Guilielmofloweria'' is an extinct genus of proterotheriid litoptern that lived from the Middle to Late Eocene of what is now Argentina. Fossils have been found in the Sarmiento Formation of Argentina. Taxonomy ''Guilielmofloweria'' was first named in 1901 by Florentino Ameghino based on fragmentary remains found in the Sarmiento Formation of Argentina, in rocks dating back to the Middle Eocene. He originally considered it a member of the pantodont family Pantolambdidae, but recent studies have shown this to be incorrect, as Richard Cifelli considered it as a member of the litoptern family Proterotheriidae, more specifically within the subfamily Anisolambdinae. Anisolambdinae (also called Anisolambdidae in some studies) was proposed to unite the primitive and earlier forms '' Anisolambda'', '' Eolicaphrium'', '' Heteroglyphis'', '' Lambdaconops'', '' Paranisolambda'', '' Protheosodon'', '' Wainka'' and '' Xesmodon''. However, the phylogenetic analysis of McGrath and colleagu ...
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Epitherium
''Epitherium'' is an extinct genus of Litopterna, who belonged to the family Proterotheriidae. It lived during the Pliocene in South America. The fossils of this herbivorous ungulate were found in Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ....The Paleobiology Database


References

Proterotheriids
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