Plesiadapiforms
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Plesiadapiforms
Plesiadapiformes (" Adapid-like" or "near Adapiformes") is a group of Primates, a sister of the Dermoptera. While none of the groups normally directly assigned to this group survived, the group appears actually not to be literally extinct (in the sense of having no living descendants) as the remaining primates (the crown primates or "Euprimates") appear to be derived Plesiadapiformes, as a sister of e.g. the Carpolestidae. The term Plesiadapiformes may still be used for all primates which are not crown primates, but this usage is paraphyletic. When the crown primates are cladistically granted, it becomes an obsolete junior synonym to primates. ''Purgatorius'' is believed to be a basal Plesiadapiformes. Plesiadapiformes first appear in the fossil record between 65 and 55 million years ago, although many were extinct by the beginning of the Eocene. They may have been the first mammals to have finger nails in place of claws. In 1990, K.C. Beard attempted to link the Plesiadapiforme ...
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Purgatorius
''Purgatorius'' is a genus of seven extinct eutherian species typically believed to be the earliest example of a primate or a proto-primate, a primatomorph precursor to the Plesiadapiformes, dating to as old as 66 million years ago. The first remains (''P. unio'' and ''P. ceratops'') were reported in 1965, from what is now eastern Montana's Tullock Formation (early Paleocene, Puercan), specifically at Purgatory Hill (hence the animal's name) in deposits believed to be about 63 million years old, and at Harbicht Hill in the lower Paleocene section of the Hell Creek Formation. Both locations are in McCone County. They have also been found in the Ravenscrag Formation and widely discovered in the early Paleocene Bug Creek Group, along with leptictids. These deposits were once thought to be late Cretaceous, but it is now clear that they are Paleocene channels with time-averaged fossil assemblages. It is thought to have been rat-sized ( long and 1.3 ounces (about 37 grams)) and a diur ...
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Paleocene
The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name is a combination of the Ancient Greek ''palaiós'' meaning "old" and the Eocene Epoch (which succeeds the Paleocene), translating to "the old part of the Eocene". The epoch is bracketed by two major events in Earth's history. The K–Pg extinction event, brought on by Chicxulub impact, an asteroid impact and possibly volcanism, marked the beginning of the Paleocene and killed off 75% of living species, most famously the non-avian dinosaurs. The end of the epoch was marked by the Paleocene–Eocene Thermal Maximum (PETM), which was a major climatic event wherein about 2,500–4,500 gigatons of carbon were released into the atmosphere and ocean systems, causing a spike in global temperatures and ocean acidification. In the Pal ...
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Adapiformes
Adapiformes is a group of early primates. Adapiforms radiated throughout much of the northern continental mass (now Europe, Asia and North America), reaching as far south as northern Africa and tropical Asia. They existed from the Eocene to the Miocene epoch. Some adapiforms resembled living lemurs. Adapiforms are known from the fossil record only, and it is unclear whether they form a monophyletic or paraphyletic group. When assumed to be a clade, they are usually grouped under the "wet-nosed" taxon Strepsirrhini, which would make them more closely related to the lemurs and less so to the "dry-nosed" Haplorhini taxon that includes monkeys and apes.Callum Ross, Richard F. Kay, ''Anthropoid origins: new visions'', Springer, 2004, , p. 100 In 2009, Franzen and colleagues placed the newly described genus ''Darwinius'' in the "Adapoidea group of early primates representative of early haplorhine diversification" so that, according to these authors, the adapiforms would not be within ...
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Euarchontoglires
Euarchontoglires (synonymous with Supraprimates) is a clade and a superorder of mammals, the living members of which belong to one of the five following groups: rodents, lagomorphs, treeshrews, colugos, and primates. Evolutionary affinities within mammals The Euarchontoglires clade is based on DNA sequence analyses and retrotransposon markers that combine the clades Glires (Rodentia + Lagomorpha) and Euarchonta (Scandentia + Primates + Dermoptera). It is usually discussed without a taxonomic rank but has been called a cohort, magnorder, or superorder. Relations among the four cohorts (Euarchontoglires, Xenarthra, Laurasiatheria, Afrotheria) and the identity of the placental root remain controversial. So far, few, if any, distinctive anatomical features have been recognized that support Euarchontoglires; nor does any strong evidence from anatomy support alternative hypotheses. Although both Euarchontoglires and diprotodont marsupials are documented to possess a vermiform appen ...
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Cladistics
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Plesiadapoidea
Plesiadapoidea was an extinct superfamily of primates that existed during the Paleocene and Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ... in Canada, Europe, and Asia.Thewissen J.G.M., Williams, E.M., and Hussain, S.T. (2001). "Eocene mammal faunas from northern Indo-Pakistan". ''Journal of Vertebrate Paleontology'' 21 (2): 347–366. DOIbr>10.1671/0272-4634(2001)021[0347:EMFFNI]2.0.CO;2Fox, Richard C.; The dentition and relationships of Carpodaptes cygneus (Russell) (Carpolestidae, Plesiadapiformes, Mammalia), from the late Paleocene of Alberta, Canada/ref> References

Plesiadapiformes Paleocene mammals Eocene mammals Paleocene first appearances {{paleo-mammal-stub ...
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Public Broadcasting Service
The Public Broadcasting Service (PBS) is an American public broadcaster and non-commercial, free-to-air television network based in Arlington, Virginia. PBS is a publicly funded nonprofit organization and the most prominent provider of educational programming to public television stations in the United States, distributing shows such as ''Frontline'', '' Nova'', ''PBS NewsHour'', ''Sesame Street'', and ''This Old House''. PBS is funded by a combination of member station dues, the Corporation for Public Broadcasting, pledge drives, and donations from both private foundations and individual citizens. All proposed funding for programming is subject to a set of standards to ensure the program is free of influence from the funding source. PBS has over 350 member television stations, many owned by educational institutions, nonprofit groups both independent or affiliated with one particular local public school district or collegiate educational institution, or entities owned by or r ...
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Claw
A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus for gripping a surface as they walk. The pincers of crabs, lobsters and scorpions, more formally known as their chelae, are sometimes called claws. A true claw is made of a hard protein called keratin. Claws are used to catch and hold prey in carnivorous mammals such as cats and dogs, but may also be used for such purposes as digging, climbing trees, self-defense and grooming, in those and other species. Similar appendages that are flat and do not come to a sharp point are called nails instead. Claw-like projections that do not form at the end of digits but spring from other parts of the foot are properly named spurs. Tetrapods In tetrapods, claws are made of keratin and consist of two layers. The unguis is the harder external layer, ...
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Finger Nails
A nail is a claw-like plate found at the tip of the fingers and toes on most primates. Nails correspond to the claws found in other animals. Fingernails and toenails are made of a tough protective protein called alpha-keratin, which is a polymer. Alpha-keratin is found in the hooves, claws, and horns of vertebrates. Structure The nail consists of the nail plate, the nail matrix and the nail bed below it, and the grooves surrounding it. Parts of the nail The matrix, sometimes called the ''matrix unguis'', keratogenous membrane, nail matrix, or onychostroma, is the active tissue (or germinal matrix) that generates cells, which harden as they move outward from the nail root to the nail plate. It is the part of the nail bed that is beneath the nail and contains nerves, lymph and blood vessels. The matrix produces cells that become the nail plate. The width and thickness of the nail plate is determined by the size, length, and thickness of the matrix, while the shape of the finger ...
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Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope Carbon-13, 13C in the atmosphere was exceptionally low in comparison with the more common isotope Carbon-12, 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Popigai impact structure, Siberia and in what is now ...
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Cladistically
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics (synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. Radi ...
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Paraphyly
In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia ( reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor, including all extant reptiles as well as the extinct synapsids, ...
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