Plasmopara Halstedii
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Plasmopara Halstedii
''Plasmopara halstedii'' is a plant pathogen infecting sunflowers. The species is one of many pathogens commonly referred to as downy mildew. ''P. halstedii'' originated in North America. ''Plasmopara halstedii'' oospores produce a thin wall which are resistant structures, sexually produced that are essential for its continuation. After entering an area, the eradication of the pathogen is difficult due to the formation of oospores, which can remain viable in soil for many years. Hosts and symptoms ''Plasmopara halstedii'' is an obligate biotroph that attacks the flowering plants of the family Asteraceae, found to infect the genus’ ''Helianthus, Bidens, Artemisia, and Xanthium.'' The pathogen has the strongest impact on ''Helianthus,'' degrading flower yields in the species '' H. argophyllus, H. debilis, H. petiolaris and H. annuus''. ''H. annuus'', the common sunflower, is also the most common host of ''P. halstedii.'' '' Xanthium strumarium'', the common cocklebur, and ' ...
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Sunflower
The common sunflower (''Helianthus annuus'') is a large annual forb of the genus ''Helianthus'' grown as a crop for its edible oily seeds. Apart from cooking oil production, it is also used as livestock forage (as a meal or a silage plant), as bird food, in some industrial applications, and as an ornamental in domestic gardens. Wild ''H. annuus'' is a widely branched annual plant with many flower heads. The domestic sunflower, however, often possesses only a single large inflorescence (flower head) atop an unbranched stem. The binomial name ''Helianthus annuus'' is derived from the Greek ''Helios'' 'sun' and ''anthos'' 'flower', while the epithet ''annuus'' means 'annual' in Latin. The plant was first domesticated in the Americas. Sunflower seeds were brought to Europe from the Americas in the 16th century, where, along with sunflower oil, they became a widespread cooking ingredient. With time, bulk of industrial-scale production has shifted to Eastern Europe, and () Russ ...
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Downy Mildew
Downy mildew refers to any of several types of oomycete microbes that are obligate parasites of plants. Downy mildews exclusively belong to the Peronosporaceae family. In commercial agriculture, they are a particular problem for growers of crucifers, grapes and vegetables that grow on vines. The prime example is ''Peronospora farinosa'' featured in NCBI-Taxonomy and HYP3. This pathogen does not produce survival structures in the northern states of the United States, and overwinters as live mildew colonies in Gulf Coast states. It progresses northward with cucurbit production each spring. Yield loss associated with downy mildew is most likely related to soft rots that occur after plant canopies collapse and sunburn occurs on fruit. Cucurbit downy mildew only affects leaves of cucurbit plants. Symptoms Initial symptoms include large, angular or blocky, yellow areas visible on the upper surface. As lesions mature, they expand rapidly and turn brown. The under surface of infected ...
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Antheridium
An antheridium is a haploid structure or organ producing and containing male gametes (called ''antherozoids'' or sperm). The plural form is antheridia, and a structure containing one or more antheridia is called an androecium. Androecium is also the collective term for the stamens of flowering plants. Antheridia are present in the gametophyte phase of cryptogams like bryophytes and ferns. Many algae and some fungi, for example ascomycetes and water moulds, also have antheridia during their reproductive stages. In gymnosperms and angiosperms, the male gametophytes have been reduced to pollen grains and in most of these the antheridia have been reduced to a single generative cell within the pollen grain. During pollination, this generative cell divides and gives rise to sperm cells. The female counterpart to the antheridium in cryptogams is the archegonium, and in flowering plants is the gynoecium. An antheridium typically consists of sterile cells and spermatogenous tissu ...
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Oogonium
An oogonium (plural oogonia) is a small diploid cell which, upon maturation, forms a primordial follicle in a female fetus or the female (haploid or diploid) gametangium of certain thallophytes. In the mammalian fetus Oogonia are formed in large numbers by mitosis early in fetal development from primordial germ cells. In humans they start to develop between weeks 4 and 8 and are present in the fetus between weeks 5 and 30. Structure Normal oogonia in human ovaries are spherical or ovoid in shape and are found amongst neighboring somatic cells and oocytes at different phases of development. Oogonia can be distinguished from neighboring somatic cells, under an electron microscope, by observing their nuclei. Oogonial nuclei contain randomly dispersed fibrillar and granular material whereas the somatic cells have a more condensed nucleus that creates a darker outline under the microscope. Oogonial nuclei also contain dense prominent nucleoli. The chromosomal material in the nucleu ...
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Heterothallism
Heterothallic species have sexes that reside in different individuals. The term is applied particularly to distinguish heterothallic fungi, which require two compatible partners to produce sexual spores, from homothallic ones, which are capable of sexual reproduction from a single organism. In heterothallic fungi, two different individuals contribute nuclei to form a zygote. Examples of heterothallism are included for ''Saccharomyces cerevisiae, Aspergillus fumigatus, Aspergillus flavus'', ''Penicillium marneffei'' and ''Neurospora crassa''. The heterothallic life cycle of ''N. crassa'' is given in some detail, since similar life cycles are present in other heterothallic fungi. Life cycle of ''Saccharomyces cerevisiae'' The yeast ''Saccharomyces cerevisiae'' is heterothallic. This means that each yeast cell is of a certain mating type and can only mate with a cell of the other mating type. During vegetative growth that ordinarily occurs when nutrients are abundant, ''S. cer ...
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Homothallism
Homothallic refers to the possession, within a single organism, of the resources to reproduce sexually; i.e., having male and female reproductive structures on the same thallus. The opposite sexual functions are performed by different cells of a single mycelium. It can be contrasted to heterothallic. It is often used to categorize fungi. In yeast, heterothallic cells have mating types a and α. An experienced mother cell (one that has divided at least once) will switch mating type every cell division cycle because of the ''HO'' allele. Sexual reproduction commonly occurs in two fundamentally different ways in fungi. These are outcrossing (in heterothallic fungi) in which two different individuals contribute nuclei to form a zygote, and self-fertilization or selfing (in homothallic fungi) in which both nuclei are derived from the same individual. Homothallism in fungi can be defined as the capability of an individual spore to produce a sexually reproducing colony when propagated ...
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Zoosporangia
A zoospore is a motility, motile asexual spore that uses a flagellum for motion (physics), locomotion. Also called a swarm spore, these spores are created by some Protist, protists, bacteria, and fungi to propagate themselves. Diversity Flagella types Zoospores may possess one or more distinct types of flagella - tinsel or "decorated", and whiplash, in various combinations. *Tinsellated (straminipilous) flagella have lateral Protein filament, filaments known as mastigonemes perpendicular to their main axis, which allow for more surface area, and disturbance of the medium, giving them the property of a rudder, that is, used for steering. *Whiplash flagella are straight, to power the zoospore through its medium. Also, the "default" zoospore only has the propelling, whiplash flagella. Both tinsel and whiplash flagella beat in a sinusoidal wave pattern, but when both are present, the tinsel beats in the opposite direction of the whiplash, to give two axes of control of motility. Mor ...
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Stoma
In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bordered by a pair of specialized parenchyma cells known as guard cells that are responsible for regulating the size of the stomatal opening. The term is usually used collectively to refer to the entire stomatal complex, consisting of the paired guard cells and the pore itself, which is referred to as the stomatal aperture. Air, containing oxygen, which is used in respiration, and carbon dioxide, which is used in photosynthesis, passes through stomata by gaseous diffusion. Water vapour diffuses through the stomata into the atmosphere in a process called transpiration. Stomata are present in the sporophyte generation of all land plant groups except liverworts. In vascular plants the number, size and distribution of stomata varies widely. ...
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Haustorium
In botany and mycology, a haustorium (plural haustoria) is a rootlike structure that grows into or around another structure to absorb water or nutrients. For example, in mistletoe or members of the broomrape family, the structure penetrates the host's tissue and draws nutrients from it. In mycology, it refers to the appendage or portion of a parasitic fungus (the hyphal tip), which performs a similar function. Microscopic haustoria penetrate the host plant's cell wall and siphon nutrients from the space between the cell wall and plasma membrane but do not penetrate the membrane itself. Larger (usually botanical, not fungal) haustoria do this at the tissue level. The etymology of the name corresponds to the Latin word ''haustor'' meaning ''the one who draws, drains or drinks'', and refers to the action performed by the outgrowth. In fungi Fungi in all major divisions form haustoria. Haustoria take several forms. Generally, on penetration, the fungus increases the surface ar ...
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Hypha
A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one or more cells surrounded by a tubular cell wall. In most fungi, hyphae are divided into cells by internal cross-walls called "septa" (singular septum). Septa are usually perforated by pores large enough for ribosomes, mitochondria, and sometimes nuclei to flow between cells. The major structural polymer in fungal cell walls is typically chitin, in contrast to plants and oomycetes that have cellulosic cell walls. Some fungi have aseptate hyphae, meaning their hyphae are not partitioned by septa. Hyphae have an average diameter of 4–6 µm. Growth Hyphae grow at their tips. During tip growth, cell walls are extended by the external assembly and polymerization of cell wall components, and the internal production of new cell membrane. The S ...
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Appressorium
An appressorium is a specialized cell typical of many fungal plant pathogens that is used to infect host plants. It is a flattened, hyphal "pressing" organ, from which a minute infection peg grows and enters the host, using turgor pressure capable of punching through even Mylar. Following spore attachment and germination on the host surface, the emerging germ tube perceives physical cues such as surface hardness and hydrophobicity, as well as chemical signals including wax monomers that trigger appressorium formation. Appressorium formation begins when the tip of the germ tube ceases polar growth, hooks, and begins to swell. The contents of the spore are then mobilized into the developing appressorium, a septum develops at the neck of the appressorium, and the germ tube and spore collapse and die. As the appressorium matures, it becomes firmly attached to the plant surface and a dense layer of melanin is laid down in the appressorium wall, except across a pore at the plant interface ...
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Zoospore
A zoospore is a motile asexual spore that uses a flagellum for locomotion. Also called a swarm spore, these spores are created by some protists, bacteria, and fungi to propagate themselves. Diversity Flagella types Zoospores may possess one or more distinct types of flagella - tinsel or "decorated", and whiplash, in various combinations. *Tinsellated (straminipilous) flagella have lateral filaments known as mastigonemes perpendicular to their main axis, which allow for more surface area, and disturbance of the medium, giving them the property of a rudder, that is, used for steering. *Whiplash flagella are straight, to power the zoospore through its medium. Also, the "default" zoospore only has the propelling, whiplash flagella. Both tinsel and whiplash flagella beat in a sinusoidal wave pattern, but when both are present, the tinsel beats in the opposite direction of the whiplash, to give two axes of control of motility. Morphological types In eukaryotes, the four main types ...
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