Pilosans Of The Caribbean
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Pilosans Of The Caribbean
The mammalian order Pilosa, which includes the sloths and anteaters, includes various species from the Caribbean region. Many species of sloths are known from the Greater Antilles, all of which became extinct over the last millennia, but some sloths and anteaters survive on islands closer to the mainland. For the purposes of this article, the "Caribbean" includes all islands in the Caribbean Sea (except for small islets close to the mainland) and the Bahamas, Turks and Caicos Islands, and Barbados, which are not in the Caribbean Sea but biogeographically belong to the same Caribbean bioregion. Overview Extinct sloths are known from the three Greater Antilles of Cuba, Hispaniola, and Puerto Rico and several smaller Antillean islands, but they are missing from the fourth of the Greater Antilles, Jamaica. These were formerly believed, on the basis of morphological studies, to be part of the family Megalonychidae, which includes some of the extinct giant ground sloths, such as ''Mega ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saur ...
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Megalonyx
''Megalonyx'' (Greek, "large claw") is an extinct genus of ground sloths of the family Megalonychidae, native to North America during the Pliocene and Pleistocene epochs. It became extinct during the Quaternary extinction event at the end of the Rancholabrean of the Pleistocene, living from ~5 million to 11,000 years ago. The type species, ''M. jeffersonii'', measured about and weighed up to . ''Megalonyx'' is descended from ''Pliometanastes,'' a genus of ground sloth that had arrived in North America during the Late Miocene, prior to the Great American Biotic Interchange. ''Megalonyx'' had the widest distribution of any North American ground sloth, having a range encompassing most of the contiguous United States, extending as far north as Alaska during warm periods. Taxonomy In 1796, Colonel John Stuart sent Thomas Jefferson, shortly before he took office as Vice President of the United States, some fossil bones: a femur fragment, ulna, radius, and foot bones including three ...
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Tamandua
''Tamandua'' is a genus of anteaters with two species: the southern tamandua (''T. tetradactyla'') and the northern tamandua (''T. mexicana''). They live in forests and grasslands, are semiarboreal, and possess partially prehensile tails. They mainly eat ants and termites, but they occasionally eat bees, beetles, and insect larvae. In captivity, they will eat fruits and meat. They have no teeth and depend on their powerful gizzards to break down their food. The word Tamanduá is Tupi-Guaraní. Extant species Description Tamanduas have tapered heads with a long, tubular snout, small eyes, and protruding ears. Their tapered mouths house a tongue reaching upwards of in length. The tail is hairless and pink in color, marked with an irregular pattern of black blotches. The forefeet possess four clawed digits, the third digit bearing the largest claw, while the hind feet have five digits. Their fur is thick, bristly, yellowish-white to fawn in color, often with a broad black lat ...
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Three-toed Sloth
The three-toed or three-fingered sloths are arboreal neotropical mammals . They are the only members of the genus ''Bradypus'' and the family Bradypodidae. The four living species of three-toed sloths are the brown-throated sloth, the maned sloth, the pale-throated sloth, and the pygmy three-toed sloth. In complete contrast to past morphological studies, which tended to place ''Bradypus'' as the sister group to all other folivorans, molecular studies place them nested within the sloth superfamily Megatherioidea, making them the only surviving members of that radiation. Extant species Evolution A study of mitochondrial cytochrome b and 16S rRNA sequences suggests that '' B. torquatus'' diverged from '' B. variegatus'' and '' B. tridactylus'' about 12 million years ago, while the latter two split 5 to 6 million years ago. The diversification of ''B. variegatus'' lineages was estimated to have started 4 to 5 million years ago. Relation to the two-toed sloth Both types of sloth t ...
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Monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have tak ...
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Biological Dispersal
Biological dispersal refers to both the movement of individuals (animals, plants, fungi, bacteria, etc.) from their birth site to their breeding site ('natal dispersal'), as well as the movement from one breeding site to another ('breeding dispersal'). Dispersal is also used to describe the movement of propagules such as seeds and spores. Technically, dispersal is defined as any movement that has the potential to lead to gene flow. The act of dispersal involves three phases: departure, transfer, settlement and there are different fitness costs and benefits associated with each of these phases. Through simply moving from one habitat patch to another, the dispersal of an individual has consequences not only for individual fitness, but also for population dynamics, population genetics, and species distribution. Understanding dispersal and the consequences both for evolutionary strategies at a species level, and for processes at an ecosystem level, requires understanding on th ...
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Oligocene
The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from the Ancient Greek (''olígos'', "few") and (''kainós'', "new"), and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period. The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion o ...
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Arboreal Locomotion
Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose numerous mechanical challenges to animals moving through them and lead to a variety of anatomical, behavioral and ecological consequences as well as variations throughout different species.Cartmill, M. (1985). Climbing. In ''Functional Vertebrate Morphology'', eds. M. Hildebrand D. M. Bramble K. F. Liem and D. B. Wake, pp. 73–88. Cambridge: Belknap Press. Furthermore, many of these same principles may be applied to climbing without trees, such as on rock piles or mountains. Some animals are exclusively arboreal in habitat, such as the tree snail. Biomechanics Arboreal habitats pose numerous mechanical challenges to animals moving in them, which have been solved in diverse ways. These challenges include moving on narrow branches, mov ...
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Radiocarbon Dating
Radiocarbon dating (also referred to as carbon dating or carbon-14 dating) is a method for determining the age of an object containing organic material by using the properties of radiocarbon, a radioactive isotope of carbon. The method was developed in the late 1940s at the University of Chicago by Willard Libby. It is based on the fact that radiocarbon () is constantly being created in the Earth's atmosphere by the interaction of cosmic rays with atmospheric nitrogen. The resulting combines with atmospheric oxygen to form radioactive carbon dioxide, which is incorporated into plants by photosynthesis; animals then acquire by eating the plants. When the animal or plant dies, it stops exchanging carbon with its environment, and thereafter the amount of it contains begins to decrease as the undergoes radioactive decay. Measuring the amount of in a sample from a dead plant or animal, such as a piece of wood or a fragment of bone, provides information that can be used to calc ...
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Megalocnidae
Megalocnidae is an extinct family of sloths, native to the islands of the Greater Antilles from the Early Oligocene to the Mid-Holocene. They are known from Cuba, Hispaniola and Puerto Rico, but are absent from Jamaica. While they were formerly placed in the Megalonychidae alongside two-toed sloths and ground sloths like ''Megalonyx'', recent mitochondrial DNA and collagen sequencing studies place them as the earliest diverging group basal to all other sloths. They displayed significant diversity in body size and lifestyle, with ''Megalocnus'' being terrestrial and probably weighing several hundred kilograms, while ''Neocnus'' was likely arboreal and similar in weight to extant tree sloths, at less than 10 kilograms. Origin It is thought that sloths arrived in the Caribbean from South America (where they arose) around the Eocene-Oligocene boundary about 33 million years ago, when there was a significant sea level drop caused by a glaciation episode. This has been associate ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Polyphyletic
A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of convergent evolution. The arrangement of the members of a polyphyletic group is called a polyphyly .. ource for pronunciation./ref> It is contrasted with monophyly and paraphyly. For example, the biological characteristic of warm-bloodedness evolved separately in the ancestors of mammals and the ancestors of birds; "warm-blooded animals" is therefore a polyphyletic grouping. Other examples of polyphyletic groups are algae, C4 photosynthetic plants, and edentates. Many taxonomists aim to avoid homoplasies in grouping taxa together, with a goal to identify and eliminate groups that are found to be polyphyletic. This is often the stimulus for major revisions of the classification schemes. Researchers concerned more with ecology than with systema ...
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