Phytosaurus
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Phytosaurus
''Phytosaurus'' (meaning "plant lizard") is a nomen dubium, dubious genus of Extinction, extinct Parasuchidae, parasuchid phytosaur found in an outcrop of the Keuper (likely the Exter Formation) in Germany. ''Phytosaurus'' was the first phytosaur to be described, being done so by Georg Friedrich von Jaeger in 1828.G. F. Jaeger. (1828). Über die Fossile Reptilien, Welche in Württemberg Aufgefunden Sind [On the Fossil Reptiles That Are Found in Württemberg]. ''Verlag der Metzler'schen Buchhandlung, Stuttgart'' 1-48 The type species is ''P. cylindricodon'' and a second species, ''P. cubicodon'', is also known. Discovery and naming In 1826, the holotype, holotypes of both species were discovered in Württemberg, Wurttemburg, Germany at the "Neckar" site at the base of the hill which Wildenau Castle stands upon. The holotype of ''P. cylindricodon'' consists of parts of the skull and jaws, with natural casts of the teeth which, however, did not preserve their conical form but were ...
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Exter Formation
The Exter Formation is the only formation of the Upper Keuper or Rhätsandstein, and is a Formation (geology), geologic formation in Germany. It preserves fossils dating back to the Rhaetian of the Triassic period (specifically around 205 Ma).Exter Formation
at Fossilworks.org
Rhätkeuper
at Fossilworks.org


Fossil content

The formation has provided fossils of:


Mammals

* ''Thomasia, Thomasia antiqua'' * Haramiyidae, Haramiyidae indet.


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Phytosaurs
Phytosaurs (Φυτόσαυροι in greek) are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. Phytosauria and Phytosauridae are often considered to be equivalent groupings containing the same species, but some studies have identified non-phytosaurid phytosaurians. Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution. The name "phytosaur" means "plant reptile", as the first fossils of phytosaurs were mistakenly thought to belong to plant eaters. The name is misleading because the sharp teeth in phytosaur jaws clearly show that they were predators. For many years, phytosaurs were considered to be the most basal group of Pseudosuchia (crocodile-line archosaurs), meaning that they were thought to be more closely related to the crocodilians than to birds (the other ...
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Phytosaur
Phytosaurs (Φυτόσαυροι in greek) are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. Phytosauria and Phytosauridae are often considered to be equivalent groupings containing the same species, but some studies have identified non-phytosaurid phytosaurians. Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution. The name "phytosaur" means "plant reptile", as the first fossils of phytosaurs were mistakenly thought to belong to plant eaters. The name is misleading because the sharp teeth in phytosaur jaws clearly show that they were predators. For many years, phytosaurs were considered to be the most basal group of Pseudosuchia (crocodile-line archosaurs), meaning that they were thought to be more closely related to the crocodilians than to birds (the othe ...
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Phytosaur
Phytosaurs (Φυτόσαυροι in greek) are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. Phytosauria and Phytosauridae are often considered to be equivalent groupings containing the same species, but some studies have identified non-phytosaurid phytosaurians. Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution. The name "phytosaur" means "plant reptile", as the first fossils of phytosaurs were mistakenly thought to belong to plant eaters. The name is misleading because the sharp teeth in phytosaur jaws clearly show that they were predators. For many years, phytosaurs were considered to be the most basal group of Pseudosuchia (crocodile-line archosaurs), meaning that they were thought to be more closely related to the crocodilians than to birds (the othe ...
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Parasuchidae
Parasuchidae is a clade of phytosaurs more derived than ''Diandongosuchus'', a basal phytosaur. It encompasses nearly all phytosaurs, include early ''Parasuchus''-grade forms as well as a more restricted clade of more specialized phytosaurs. This more restricted clade is traditionally known as the family Phytosauridae and more recently as the subfamily Mystriosuchinae. Parasuchids have been recovered from Late Triassic deposits in Europe, North America, India, Morocco, Thailand, Brazil, Greenland and Madagascar. In their osteology of ''Parasuchus'', Kammerer et al. (2016) suggested using Parasuchidae to include taxa traditionally included in Phytosauridae as well as ''Parasuchus''-grade taxa. Stocker et al. (2017) use the phytosaur classification advocated by Kammerer et al. (2016) by recovering ''Diandongosuchus ''Diandongosuchus'' is an extinct genus of archosauriform reptile, possibly a member of the Phytosauria, known from the Middle Triassic of China. The type species '' ...
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Mystriosuchinae
Parasuchidae is a clade of phytosaurs more derived than ''Diandongosuchus'', a basal phytosaur. It encompasses nearly all phytosaurs, include early ''Parasuchus''-grade forms as well as a more restricted clade of more specialized phytosaurs. This more restricted clade is traditionally known as the family Phytosauridae and more recently as the subfamily Mystriosuchinae. Parasuchids have been recovered from Late Triassic deposits in Europe, North America, India, Morocco, Thailand, Brazil, Greenland and Madagascar. In their osteology of ''Parasuchus'', Kammerer et al. (2016) suggested using Parasuchidae to include taxa traditionally included in Phytosauridae as well as ''Parasuchus''-grade taxa. Stocker et al. (2017) use the phytosaur classification advocated by Kammerer et al. (2016) by recovering ''Diandongosuchus ''Diandongosuchus'' is an extinct genus of archosauriform reptile, possibly a member of the Phytosauria, known from the Middle Triassic of China. The type species ' ...
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Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', t ...
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Nomen Dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a specimen belongs to that group or not. This may happen if the original type series (i. e. holotype, isotype, syntype or paratype) is lost or destroyed. The zoological and botanical codes allow for a new type specimen, or neotype, to be chosen in this case. A name may also be considered a ''nomen dubium'' if its name-bearing type is fragmentary or lacking important diagnostic features (this is often the case for species known only as fossils). To preserve stability of names, the ''International Code of Zoological Nomenclature'' allows a new type specimen, or neotype, to be chosen for a ''nomen dubium'' in this case. 75.5. Replacement of unidentifiable name-bearing type by a neotype. When an author considers that the taxonomic identity of a ...
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Nomina Dubia
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a specimen belongs to that group or not. This may happen if the original type series (i. e. holotype, isotype, syntype or paratype) is lost or destroyed. The zoological and botanical codes allow for a new type specimen, or neotype, to be chosen in this case. A name may also be considered a ''nomen dubium'' if its name-bearing type is fragmentary or lacking important diagnostic features (this is often the case for species known only as fossils). To preserve stability of names, the ''International Code of Zoological Nomenclature'' allows a new type specimen, or neotype, to be chosen for a ''nomen dubium'' in this case. 75.5. Replacement of unidentifiable name-bearing type by a neotype. When an author considers that the taxonomic identity of a ...
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Texas
Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by both List of U.S. states and territories by area, area (after Alaska) and List of U.S. states and territories by population, population (after California). Texas shares borders with the states of Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the west, and the Mexico, Mexican States of Mexico, states of Chihuahua (state), Chihuahua, Coahuila, Nuevo León, and Tamaulipas to the south and southwest; and has a coastline with the Gulf of Mexico to the southeast. Houston is the List of cities in Texas by population, most populous city in Texas and the List of United States cities by population, fourth-largest in the U.S., while San Antonio is the second most pop ...
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Dockum Formation
The Dockum is a Late Triassic (approximately late Carnian through Rhaetian, or 223–200 Ma) geologic group found primarily on the Llano Estacado of western Texas and eastern New Mexico with minor exposures in southwestern Kansas, eastern Colorado, and Oklahoma panhandle. The Dockum reaches a maximum thickness of slightly over 650 m but is usually much thinner. The Dockum rests on an unconformity over the Anisian (242–234 Ma) aged Anton Chico Formation. The Dockum and Chinle Formation were deposited roughly at the same time and share many of the same vertebrates and plant fossils. They appear to have very similar paleoenviroments. The two units are approximately separated by the Rio Grande in central New Mexico. This has led to controversy over the stratigraphic nomenclature for the Chinle and Dockum. History of investigation There is no designated type locality for this formation. The Dockum was named by William Fletcher Cummins for a good exposure in vicinity of town ...
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Hans-Dieter Sues
Hans-Dieter Sues (born January 13, 1956) is a German-born American paleontologist who is Senior Scientist and Curator of Vertebrate Paleontology at the National Museum of Natural History of the Smithsonian Institution in Washington, DC. He received his education at Johannes Gutenberg-Universität Mainz (University of Mainz), University of Alberta, and Harvard University (Ph.D., 1984). Before assuming his present position, Sues worked at the Royal Ontario Museum and the University of Toronto and at the Carnegie Museum of Natural History in Pittsburgh. He is interested in the diversity, paleoecology, and evolutionary history of Paleozoic and Mesozoic tetrapods, especially archosaurian reptiles and cynodont therapsids, and the history of biology and paleontology. Sues has discovered numerous new dinosaurs and other extinct terrestrial vertebrates in Paleozoic and Mesozoic continental strata in North America and Europe. He has authored or co-authored over 150 articles and book chapte ...
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