Physornis
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Physornis
''Physornis'' is an extinct genus of giant flightless predatory birds of the family Phorusrhacidae or "terror birds", most closely related to '' Paraphysornis'', that lived in Argentina. The type species is ''P. fortis''. It lived during the Middle to Late Oligocene ( Deseadan). Few fossils are known, but the available material suggests that ''Physornis'' was one of the largest phorusrhacids. History and taxonomy The holotype of ''Physornis'' ( BMNH-A583) is a 137 mm long portion of a symphysis and the right branch of this lower jaw, described in 1894 by paleontologist Florentino Ameghino. He published the name, together with a description but without a drawing, in the same year.Brodkorb, P. (1967). ''Catalogue of fossil birds: part 3 (Ralliformes, Ichthyornithiformes, Charadriiformes)''. University of Florida.Alvarenga, H. M., & Höfling, E. (2003). Systematic revision of the Phorusrhacidae (Aves: Ralliformes). ''Papéis Avulsos de Zoologia'', ''43''(4), 55-91. The fossil was ...
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Paraphysornis
''Paraphysornis'' is an extinct genus of giant flightless terror birds that inhabited Brazil during Late Oligocene or Early Miocene epochs. Although not the tallest phorusrhacid, ''Paraphysornis'' was a notably robust bird, having short and robust tarsal bones not suited for pursuit hunting. Discovery and naming The holotype (DGM-1418-R) and only known fossil was discovered by a mining company in the Taubaté Basin (São Paulo State) in a layer of bentonite clay, two or three meters below a layer of Pyrobitumen. These sediments belong to the Late Oligocene to Early Miocene Tremembé Formation. Excavation of the bones took several months between 1977 and 1978, yielding a nearly complete (%75) skeleton only lacking most of the cranium, the pelvis and sternum. The bones were initially studied by Herculano Alvarenga, who first described them in 1982 as a species of ''Physornis''. Following closer examination of various phorusrhacid remains held in institutions across Europe, Nor ...
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Patagornis
''Patagornis'' is a genus of extinct flightless predatory birds of the family Phorusrhacidae. Known as "terror birds", these lived in what is now Argentina during the Early and Middle Miocene; the Santa Cruz Formation in Patagonia contains numerous specimens.''Patagornis''
at Fossilworks.org
''Patagornis'' was an agile, medium sized Patagornithine and was likely a pursuit predator.


Etymology

The genus name means "bird from Patagonia" after the location the fossils were found in and the species name after prominent American paleontologist ...
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Brontornis
''Brontornis'' is an extinct genus of giant bird that inhabited Argentina during the Early to Middle Miocene. Its taxonomic position is highly controversial, with authors alternatively considering it to be a cariamiform, typically a phorusrhacid (terror bird) or an anserimorph. Taxonomy The first fossils of ''Brontornis burmeisteri'' were described by paleontologists Francisco Moreno and Alcides Mercerart in 1891, the fossils being a left femur, tibiotarsus, fibula, and tarsometatarsus all from the same individual found in the Lower-Middle Miocene strata of the Santa Cruz Formation in Santa Cruz Province, Argentina.Brodkorb, P. (1967). ''Catalogue of fossil birds: part 3 (Ralliformes, Ichthyornithiformes, Charadriiformes)''. University of Florida. In the same paper, two distal tibiotarsi from the same area were referred to ''Brontornis'' as well.Moreno, F. P., & Mercerat, A. (1891). ''Catálogo de los pájaros fósiles de la República Argentina conservados en el Museo de La ...
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Andrewsornis
''Andrewsornis'' is an extinct genus of giant flightless predatory birds of the family Phorusrhacidae or "terror birds" that lived in Oligocene Argentina. Fossils have been found in the Sarmiento Formation, and possibly the Agua de la Piedra Formation.''Andrewsornis''
at Fossilworks.org


Discovery and naming

The of ''Andrewsornis'' ( FM-P13417) was discovered on September 18, 1923 by John Bernard Abbott in

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Deseadan
The Deseadan ( es, Deseadense) age is a period of geologic time (29.0–21.0 Ma) within the Oligocene epoch of the Paleogene to the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification of South America. It follows the Tinguirirican and precedes the Colhuehuapian age. Etymology The age is named after the Deseado Formation of the Deseado Massif in eastern Patagonia, Argentina. Formations Fossils Correlations The Deseadan South American land mammal age (SALMA) is equivalent to the Arikareean in the North American land mammal age (NALMA) and the Harrisonian The Harrisonian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 24,800,000 to 20,600,000 years BP, a period of . It is usual ... in the 2000 version of the classification. It overlaps with the Hsandagolian of Asia and the MP 25 zone of Europe, the Waitak ...
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Phorusrhacidae
Phorusrhacids, colloquially known as terror birds, are an extinct clade of large carnivorous flightless birds that were one of the largest species of apex predators in South America during the Cenozoic era; their conventionally accepted temporal range covers from 62 to 0.1 million years ( Ma) ago. They ranged in height from . Their closest modern-day relatives are believed to be the seriemas. '' Titanis walleri'', one of the larger species, is known from Texas and Florida in North America. This makes the phorusrhacids the only known large South American predator to migrate north in the Great American Interchange that followed the formation of the Isthmus of Panama land bridge (the main pulse of the interchange began about 2.6 Ma ago; ''Titanis'' at 5 Ma was an early northward migrant). It was once believed that ''T. walleri'' became extinct in North America around the time of the arrival of humans, but subsequent datings of ''Titanis'' fossils provided no evidence for their surv ...
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Herculano Marcos Ferraz De Alvarenga
Herculano Marcos Ferraz de Alvarenga (born 7 November 1947) is a Brazilian ornithologist, paleontologist and physician, founder of the Taubaté Natural History Museum.Alvarenga, H. (2004) Tucanos das Américas/Toucans of the Americas. M. Pontual Ed.: 120pp.Museu de História Natural de Taubaté. (UndatedHistórico Retrieved 30 January 2017. Life Herculano Alvarenga was born in 1947 in Taubaté, São Paulo, Brazil. As a teenager, he started to watch birds and collect them. He went to São Paulo when he was 15 to study taxidermy and soon his stuffed specimens started to be exhibited in scientific expositions in high school.Moon, P. (2012).Herculano Alvarenga descobriu fósseis e hoje dirige seu próprio museu de história natural. ''Época''. Retrieved 4 February 2017. His interests in biology and anatomy led him to study medicine, specializing in orthopedy. In 1975 he returned to Taubaté and became professor of the Faculty of Medicine in the city. When the faculty went on st ...
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Species Inquirenda
In biological classification, a ''species inquirenda'' is a species of doubtful identity requiring further investigation. The use of the term in English-language biological literature dates back to at least the early nineteenth century. The term taxon inquirendum is broader in meaning and refers to an incompletely defined taxon of which the taxonomic validity is uncertain or disputed by different experts or is impossible to identify the taxon. Further characterization is required. See also * Glossary of scientific naming * ''Candidatus'', a proposed taxa based on incomplete evidence * ''incertae sedis'', a taxon of uncertain position in a classification * '' nomen dubium'', a name of unknown or doubtful application * Open nomenclature Open nomenclature is a vocabulary of partly informal terms and signs in which a taxonomist may express remarks about their own material. This is in contrast to synonymy lists, in which a taxonomist may express remarks on the work of others. Common . ...
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Nomen Dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a specimen belongs to that group or not. This may happen if the original type series (i. e. holotype, isotype, syntype or paratype) is lost or destroyed. The zoological and botanical codes allow for a new type specimen, or neotype, to be chosen in this case. A name may also be considered a ''nomen dubium'' if its name-bearing type is fragmentary or lacking important diagnostic features (this is often the case for species known only as fossils). To preserve stability of names, the ''International Code of Zoological Nomenclature'' allows a new type specimen, or neotype, to be chosen for a ''nomen dubium'' in this case. 75.5. Replacement of unidentifiable name-bearing type by a neotype. When an author considers that the taxonomic identity ...
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Upper Oligocene
The Chattian is, in the geologic timescale, the younger of two ages or upper of two stages of the Oligocene Epoch/Series. It spans the time between . The Chattian is preceded by the Rupelian and is followed by the Aquitanian (the lowest stage of the Miocene). Stratigraphic definition The Chattian was introduced by Austrian palaeontologist Theodor Fuchs in 1894. Fuchs named the stage after the Chatti, a Germanic tribe.Berry, Edward W"The Mayence Basin, a Chapter of Geologic History" ''The Scientific Monthly'', Vol. 16, No. 2, February 1923. pp. 114. Retrieved March 18, 2020. The original type locality was near the German city of Kassel. The base of the Chattian is at the extinction of the foram genus ''Chiloguembelina'' (which is also the base of foram biozone P21b). An official GSSP for the Chattian Stage was ratified in October of 2016. The top of the Chattian Stage (which is the base of the Aquitanian Stage, Miocene Series and Neogene System) is at the first appearance of ...
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Early Miocene
The Early Miocene (also known as Lower Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages: the Aquitanian age, Aquitanian and Burdigalian stages. The sub-epoch lasted from 23.03 ± 0.05 annum, Ma to 15.97 ± 0.05 Ma (million years ago). It was preceded by the Oligocene epoch. As the climate started to get cooler, the landscape started to change. New mammals evolved to replace the extinct animals of the Oligocene epoch. The first members of the hyena and weasel family started to evolve to replace the extinct ''Hyaenodon'', entelodonts and bear-dogs. The chalicotheres survived the Oligocene epoch. A new genus of entelodont called ''Daeodon'' evolved in order to adapt to the new habitats and hunt the new prey animals of the Early Miocene epoch; it quickly became the top predator of North America. But it became extinct due to competition from ''Amphicyon'', a newcomer from Eurasia. ''Amphicyon'' bested ''Daeodon'' because the bear-dog's ...
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Phalanx Bone
The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bones. Structure The phalanges are the bones that make up the fingers of the hand and the toes of the foot. There are 56 phalanges in the human body, with fourteen on each hand and foot. Three phalanges are present on each finger and toe, with the exception of the thumb and large toe, which possess only two. The middle and far phalanges of the fifth toes are often fused together (symphalangism). The phalanges of the hand are commonly known as the finger bones. The phalanges of the foot differ from the hand in that they are often shorter and more compressed, especially in the proximal phalanges, those closest to the torso. A phalanx is named according to whether it is proximal, middle, or distal and its associated finger or toe. The p ...
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