Perichelydia
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Perichelydia
Perichelydia (from Greek ''peri'' "near" and ''chelys'' "turtle") is a clade within Pantestudines (turtles and their extinct relatives) known from the Middle Jurassic to Holocene. Alongside crown group Testudines, it also contains Helochelydridae, which is known from the Cretaceous of Europe and North America, Sichuanchelyidae from the Middle Jurassic to Paleocene of Asia and Europe, Meiolaniformes, which is known from the Cretaceous to Holocene of South America, Australia and Oceania, and '' Spoochelys'', known from the Mid-Cretaceous Griman Creek Formation of Australia. ''Kallokibotion'' from the Late Cretaceous of Europe is also considered part of this group. Several other groups, including the proposed clade Angolachelonia (containing Thalassochelydia and Sandownidae), Paracryptodira, Macrobaenidae, Sinemydidae and Xinjiangchelyidae, which are sometimes considered members of Cryptodira, have also been found outside crown Testudines in several analyses. These groups are usually ...
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Mesochelydia
Mesochelydia (from Greek ''mesos'' "middle" and ''chelys'' "turtle") is a clade within Pantestudines, more inclusive than Perichelydia, but less than Testudinata. The clade is known from the Early Jurassic to the Present, and contains all Jurassic representatives of Testudinata aside from ''Australochelys.'' The ancestral condition for Mesochelydia is thought to be aquatic, as opposed to terrestrial for Testudinata. They are distinguished from more basal testudinatans by the presence of the following characters: strap like pectoral girdle, supramarginals absent, reduced posterior entoplastral process, eleven pairs of peripherals, elongate processus interfenestralis, paired basioccipital tubercles, fully formed cavum tympani and antrum postoticum, single vomer, confluent external nares, lacrimals and supratemporals absent. Taxonomy * ''Condorchelys'' Sterli, 2008 Cañadón Asfalto Formation, Middle Jurassic, Argentina * ''Eileanchelys'' Anquetin et al. 2009 Kilmaluag Formation, ...
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Helochelydridae
The Helochelydridae are an extinct family of stem-turtles known from fossils found in North America and Europe spanning the Early to Late Cretaceous. Description The skull, shell and osteoderms of helochelydrids are covered in small, cylindrical protuberances, which are a distinctive characteristic of the group. They are thought to be terrestrial, based on the presence of limb osteoderms and bone histology. Their skull morphology is dissimilar to that of extant tortoises, suggesting an omnivorous habit similar to that of box turtles. Taxonomy Helochelydridae includes all turtles that closely related to '' Helochelydra'' than '' Sichuanchelys'', '' Meiolania'', or extant turtles. Although referred to as Solemydidae in recent literature on extinct turtles, Helochelydridae has priority over Solemydidae. They are placed as part of the clade Perichelydia. Some recent studies have recovered them as paracryptodires, though other studies have found them to be more basal than parac ...
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Meiolaniformes
Meiolaniformes is an extinct clade of stem-group turtles, defined as all taxa more closely related to ''Meiolania'' than to Cryptodira and Pleurodira. It is known from the Early Cretaceous to the Holocene of Australia, Oceania and South America. Some Eurasian taxa have been suggested to be part of the group, but this is disputed. The oldest member of Meiolaniformes is Australian Early Cretaceous '' Otwayemys''. Taxonomy The clade as currently defined contains the following taxa: * †''Chubutemys'' Gaffney et al. 2007 - Aptian, Cerro Barcino Formation, Argentina * †'' Otwayemys'' Gaffney et al. 1998 - Aptian, Eumeralla Formation, Australia * †''Patagoniaemys'' Sterli and de la Fuente 2011 Upper Campanian-Lower Maastrichtian, La Colonia Formation, Argentina * †'' Peligrochelys'' Sterli and de la Fuente 2011 Danian, Salamanca Formation, Argentina * †'' Trapalcochelys'' Sterli et al. 2013 Upper Campanian, Allen Formation, Argentina * †Meiolaniidae Lydekker 1887, Eocene ...
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Sichuanchelyidae
Sichuanchelyidae is a family of extinct turtles in the clade Testudinata. It includes all perichelydians that are more closely related to '' Sichuanchelys'' than '' Meiolania'', '' Helochelydra'', or any extant turtles. Systematics The family Sichuanchelyidae was originally created to accommodate the Middle Jurassic Chinese turtle '' Sichuanchelys'', and it was considered monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ... until Joyce et al. (2016) recovered the problematic Late Cretaceous turtle '' Mongolochelys'' from Mongolia as a close relative of ''Sichuanchelys''. The late Paleocene form '' Laurasichersis'' from Europe is the youngest sichuanchelyid, showing that stem turtles in Laurasia outlived the Mesozoic. References Extinct turtles Testudinata Prehisto ...
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Griman Creek Formation
The Griman Creek Formation is a geological formation in northern New South Wales and southern Queensland, Australia whose strata date back to the Albian-Cenomanian of the Early-Late Cretaceous.Bell et al., 2019 It is most notable being a major source of opal, found near the town of Lightning Ridge, New South Wales. Alongside the opal opalised fossils are also found, including those of dinosaurs and primitive monotremes.Weishampel ''et al.'', 2004, pp.573-574 Description As a whole, the formation primarily consists of thinly bedded medium to fine sandstone, siltstone and mudstone, with sporadic coal seams. In the vicinity of Lightning Ridge, it is divided up into two informal members the underlying Wallangulla Sandstone Member which primarily consists of red fine grained sandstone, light siltstone and grey claystone and is up to thick while the overlying Coocoran Claystone consists of about 10 metres of claystone. The contact between the two units is sudden and unconformable. ...
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Pantestudines
Pantestudines or Pan-Testudines is the group of all reptiles more closely related to turtles than to any other living animal. It includes both modern turtles (crown group turtles, also known as Testudines) and all of their extinct relatives (also known as stem-turtles). Classification The identity of the ancestors and closest relatives of the turtle lineage was a longstanding scientific mystery, though new discoveries and better analyses in the early 21st century began to clarify turtle relationships. Analysis of fossil data has shown that turtles are diapsid reptiles, most closely related either to the archosaurs (crocodiles, bird, and relatives) or the lepidosaurs (lizards, tuatara, and relatives). Genetic analysis strongly favors the hypothesis that turtles are the closest relatives of the archosaurs, though studies using only fossil evidence often continue to recover them as relatives of lepidosaurs. Studies using only fossils, as well as studies using a combination of fossil ...
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Cryptodira
The Cryptodira ('' el, hidden neck'') are a suborder of Testudines that includes most living tortoises and turtles. Cryptodira differ from Pleurodira (side-necked turtles) in that they lower their necks and pull the heads straight back into the shells, instead of folding their necks sideways along the body under the shells' marginals. They include among their species freshwater turtles, snapping turtles, tortoises, softshell turtles, and sea turtles. Neck retraction The Cryptodira are characterized by retraction of the head in the vertical plane, which permits for primarily vertical movements and restricted lateral movements outside of the shell. These motions are largely due to the morphology and arrangement of cervical vertebrae. In all recent turtles, the cervical column consists of nine joints and eight vertebrae. Compared to the narrow vertebrae and the closely positioned zygapophyses of the pleurodires, the cryptodires’ vertebrae take on the opposite shape. Their ce ...
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Sandownidae
Sandownidae is a family of extinct marine turtles from the Cretaceous and Paleogene distributed around the Atlantic Ocean and adjacent areas. The family is defined as all taxa closer to the type genus '' Sandownia'' than to ''Pelomedusa'', '' Testudo'', ''Solnhofia'', ''Eurysternum'', ''Plesiochelys'', ''Thalassemys'' or '' Protostega'', a definition that encompasses the previous concept of the clade while also excluding it from being synonymous with other clades of modern or extinct marine turtles. Sandownidae may be within the larger clade Angolachelonia, defined as inclusive of ''Angolachelys'' and ''Solnhofia'', sister to the entirely Late Jurassic marine group Thalassochelydia, although the concepts of the clades may shift with further phylogenetic analysis. Genera * '' Sandownia'' Meylan et al. 2000 Atherfield Clay Formation, England, Early Cretaceous (Aptian) * '' Brachyopsemys'' Tong and Meylan 2013 Ouled Abdoun Basin, Morocco, Paleocene (Danian) * ''Angolachelys'' Mate ...
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Thalassochelydia
Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including ''Eurysternum'', ''Plesiochelys'' and ''Thalassemys'' to the exclusion of ''Pelomedusa'', '' Testudo'' and '' Protostega''. While a clade uniting the families Eurysternidae, Plesiochelyidae and Thalassemydidae had been supported by phylogenetic evidence, a name was not given for the clade until 2017, when Jérémy Anquetin and colleagues coined Thalassochelydia. While inner relationships of the clade are so uncertain as to make the monophyly of the families questionable, the significant diversity of thalassochelydians makes using the potentially invalid terms useful to describe the different morphologies. The monophyly of Thalassochelydia was further supported by a later analysis by Serjocha Evers and Roger Benson in 2019, where the group, represented by '' Solnhofia'' and Plesiochelyidae was sister taxon to Sandown ...
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Paleocene
The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name is a combination of the Ancient Greek ''palaiós'' meaning "old" and the Eocene Epoch (which succeeds the Paleocene), translating to "the old part of the Eocene". The epoch is bracketed by two major events in Earth's history. The K–Pg extinction event, brought on by Chicxulub impact, an asteroid impact and possibly volcanism, marked the beginning of the Paleocene and killed off 75% of living species, most famously the non-avian dinosaurs. The end of the epoch was marked by the Paleocene–Eocene Thermal Maximum (PETM), which was a major climatic event wherein about 2,500–4,500 gigatons of carbon were released into the atmosphere and ocean systems, causing a spike in global temperatures and ocean acidification. In the Pal ...
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Turtle
Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtles), which differ in the way the head retracts. There are 360 living and recently extinct species of turtles, including land-dwelling tortoises and freshwater terrapins. They are found on most continents, some islands and, in the case of sea turtles, much of the ocean. Like other amniotes (reptiles, birds, and mammals) they breathe air and do not lay eggs underwater, although many species live in or around water. Turtle shells are made mostly of bone; the upper part is the domed carapace, while the underside is the flatter plastron or belly-plate. Its outer surface is covered in scales made of keratin, the material of hair, horns, and claws. The carapace bones develop from ribs that grow sideways and develop into broad flat plates th ...
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Crown Group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown gr ...
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