Pentazonia
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Pentazonia
Pentazonia is a taxonomic infraclass of millipedes containing the pill-millipedes (Oniscomorpha) which can roll into a ball and the order Glomeridesmida which cannot. Defining traits (apomorphies) include divided sternites, a labrum with single median tooth, and an enlarged pygidium on the hind-most body segment. Pentazonia is in the dominant millipede subclass Chilognatha which have a calcified exoskeleton and modified sperm-transferring legs in males. In contrast to the Helminthomorpha – the other Chilognathan infraclass, the sperm-transferring legs are located on posterior body segments and known as ''telopods''. Pentazonians are relatively short-bodied, with between 13 and 21 body segments. The Pentazonia contains one extinct order, Amynilyspedida Pill millipedes are any members of two living (and one extinct) Order (biology), orders of millipedes, often grouped together into a single superorder, Oniscomorpha. The name Oniscomorpha refers to the millipedes' resemblance t ...
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Millipede
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from the Latin for "thousand feet", no species was known to have 1,000 or more until the discovery of ''Eumillipes persephone'', which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures. Most millipedes are slow-moving detritivores, eating decaying leaves and other dead plant matter. Some eat fungi or ...
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Limacomorpha
Glomeridesmida is an order of millipedes in the infraclass Pentazonia containing 2 families and at least 31 species. Glomeridesmida is the only living order of the superorder Limacomorpha. Glomeridesmidans are small (less than ) and somewhat flattened, and unlike other orders of Pentazonia, are unable to roll into a ball. Ocelli (eyes) are absent. Adult females in this order have 36 pairs of legs and 21 segments, counting 20 tergites plus the anal shield. Male specimens in this order are rare, unknown for all species in the family Termitodesmidae, and known for only a small number of species in the family Glomeridesmidae. Descriptions of mature males in at least three species (''Glomeridesmus spelaeus'', ''G. siamensis'', and ''G. marmoreus'') report 35 pairs of legs, including a pair of telopods, and 20 segments, one fewer than the 21 segments found in adult females. The description of an adult male of another species (''G. indus''), however, reports 37 pairs of legs, includ ...
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Glomeridesmida
Glomeridesmida is an order of millipedes in the infraclass Pentazonia containing 2 families and at least 31 species. Glomeridesmida is the only living order of the superorder Limacomorpha. Glomeridesmidans are small (less than ) and somewhat flattened, and unlike other orders of Pentazonia, are unable to roll into a ball. Ocelli (eyes) are absent. Adult females in this order have 36 pairs of legs and 21 segments, counting 20 tergites plus the anal shield. Male specimens in this order are rare, unknown for all species in the family Termitodesmidae, and known for only a small number of species in the family Glomeridesmidae. Descriptions of mature males in at least three species (''Glomeridesmus spelaeus'', ''G. siamensis'', and ''G. marmoreus'') report 35 pairs of legs, including a pair of telopods, and 20 segments, one fewer than the 21 segments found in adult females. The description of an adult male of another species (''G. indus''), however, reports 37 pairs of legs, includi ...
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Chilognatha
Chilognatha is a subclass of the class Diplopoda, which includes the vast majority of extant millipedes, about 12,000 species. Taxonomy The classification of Chilognatha presented below is based on Shear, 2011, and Shear & Edgecombe, 2010 (extinct groups). Recent cladistic and molecular studies have challenged the traditional classification schemes above, and in particular the position of the orders Siphoniulida and Polyzoniida is not yet well established. The placement and positions of extinct groups (†) known only from fossils is tentative and not fully resolved. After each name is listed the author citation: the name of the person who coined the name or defined the group, even if not at the current rank. * Subclass Chilognatha Latrielle, 1802 ** Order † Zosterogrammida Wilson, 2005 (Chilognatha ''incertae sedis'') ** Infraclass Pentazonia Brandt, 1833 *** Order † Amynilyspedida Hoffman, 1969 *** Superorder Limacomorpha Pocock, 1894 **** Order Glomeridesmida Cook, ...
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Helminthomorpha
Chilognatha is a subclass of the class Diplopoda, which includes the vast majority of extant millipedes, about 12,000 species. Taxonomy The classification of Chilognatha presented below is based on Shear, 2011, and Shear & Edgecombe, 2010 (extinct groups). Recent cladistic and molecular studies have challenged the traditional classification schemes above, and in particular the position of the orders Siphoniulida and Polyzoniida is not yet well established. The placement and positions of extinct groups (†) known only from fossils is tentative and not fully resolved. After each name is listed the author citation: the name of the person who coined the name or defined the group, even if not at the current rank. * Subclass Chilognatha Latrielle, 1802 ** Order † Zosterogrammida Wilson, 2005 (Chilognatha ''incertae sedis'') ** Infraclass Pentazonia Brandt, 1833 *** Order †Amynilyspedida Hoffman, 1969 *** Superorder Limacomorpha Pocock, 1894 **** Order Glomeridesmida Cook, 18 ...
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Glomeris Marginata
''Glomeris marginata'' is a common European species of pill millipede. It is a short millipede, rounded in cross-section, which is capable of rolling itself up into a ball ("volvation") when disturbed. This behaviour is also found in the pill woodlouse ''Armadillidium'', with which ''G. marginata'' is often confused. Distribution ''Glomeris marginata'' is found throughout central and north-western Europe, from Poland and Scandinavia to Spain and Italy. In the British Isles, it is found in all areas south of the Central Belt of Scotland. Description ''Glomeris marginata'' grows up to long and wide, and is covered by twelve black dorsal plates with white rims. Each segment except those at the front and back bears two pairs of legs, with around 18 pairs in total. This distinguishes pill millipedes from pill woodlice, both of which are called "pillbugs" — woodlice have 7 pairs of walking legs, one per body segment, while millipedes have more pairs, and with two pairs to each ap ...
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Johann Friedrich Von Brandt
Johann Friedrich von Brandt (25 May 1802 – 15 July 1879) was a German-Russian natural history, naturalist, who worked mostly in Russia. Brandt was born in Jüterbog and educated at a Gymnasium (school), gymnasium in Wittenberg and the Humboldt University of Berlin, University of Berlin. In 1831 he emigrated to Russia, and soon was appointed director of the Zoological Museum of the St Petersburg Academy of Sciences. Brandt encouraged the collection of native animals, many of which were not represented in the museum. Many specimens began to arrive from the expeditions of Nikolai Alekseevich Severtzov, Severtzov, Nikolai Przhevalsky, Przhevalsky, Aleksandr Fyodorovich Middendorf, Middendorff, Leopold von Schrenck, Schrenck and Gustav Radde. He described several birds collected by Russian explorers off the Pacific Coast of North America, including Brandt's cormorant, red-legged kittiwake and spectacled eider. As a paleontologist, Brandt ranks among the best. He was also an entomo ...
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Oniscomorpha
Pill millipedes are any members of two living (and one extinct) Order (biology), orders of millipedes, often grouped together into a single superorder, Oniscomorpha. The name Oniscomorpha refers to the millipedes' resemblance to certain woodlouse, woodlice (Oniscidea), also called pillbugs or "roly-polies". However, millipedes and woodlice are not closely related (belonging to the subphylum, subphyla Myriapoda and Crustacean, Crustacea, respectively); rather, this is a case of convergent evolution. Description Pill millipedes are relatively short-bodied compared to most other millipedes, with only eleven to thirteen body segments, and are capable of rolling into a ball (volvation) when disturbed, as a antipredator adaptation, defense against predators. This ability evolution, evolved separately in each of the two orders, making it a case of convergent evolution, rather than Homology (biology), homology. They can also millipede#Defence mechanisms, exude a noxious liquid, which may ...
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Pill-millipede
Pill millipedes are any members of two living (and one extinct) orders of millipedes, often grouped together into a single superorder, Oniscomorpha. The name Oniscomorpha refers to the millipedes' resemblance to certain woodlice (Oniscidea), also called pillbugs or "roly-polies". However, millipedes and woodlice are not closely related (belonging to the subphyla Myriapoda and Crustacea, respectively); rather, this is a case of convergent evolution. Description Pill millipedes are relatively short-bodied compared to most other millipedes, with only eleven to thirteen body segments, and are capable of rolling into a ball (volvation) when disturbed, as a defense against predators. This ability evolved separately in each of the two orders, making it a case of convergent evolution, rather than homology. They can also exude a noxious liquid, which may be both caustic and toxic, to repel predators. Pill millipedes are detritivorous, feeding on decomposing plant matter, usually in ...
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Apomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis Th ...
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Sternites
The sternum (pl. "sterna") is the ventral portion of a segment of an arthropod thorax or abdomen. In insects, the sterna are usually single, large sclerites, and external. However, they can sometimes be divided in two or more, in which case the subunits are called sternites, and may also be modified on the terminal abdominal segments so as to form part of the functional genitalia, in which case they are frequently reduced in size and development, and may become internalized and/or membranous. For a detailed explanation of the terminology, see Kinorhynchs have tergal and sternal plates too, though seemingly not homologous with those of arthropods.Sørensen, M. V. et al. Phylogeny of Kinorhyncha based on morphology and two molecular loci. PLoS One 10, 1–33 (2015). Ventrites are externally visible sternites. Usually the first sternite is covered up, so that vertrite numbers do not correspond to sternid numbers. The term is also used in other arthropod groups such as crustaceans ...
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Labrum (arthropod Mouthpart)
The labrum is a flap-like structure that lies immediately in front of the mouth in almost all extant Euarthropoda. The most conspicuous exceptions are the Pycnogonida, which probably are chelicerate-relatives. In entomology, the labrum amounts to the "upper lip" of an insect mouth, the corresponding "lower lip" being the labium. The evolutionary origin, embryogenesis and morphological development of the labrum have proved to be by far the most controversial and challenging topic in the study of arthropod head structures. Embryonic nature and origin of the labrum The labrum is innervated in crustaceans and insects from the tritocerebrum (the back of the brain). However, in development, its embryonic primordium often appears at the anterior of the head and migrates backwards towards its adult position. Furthermore, it often appears as a bilobed structure, with a set of muscles, nerves and gene expression in many ways similar to that of an appendage. This evidence has been use ...
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