Peloneustes Philarchus
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Peloneustes Philarchus
''Peloneustes'' (meaning "mud swimmer") is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally described as a species of ''Plesiosaurus'' by palaeontologist Harry Govier Seeley in 1896, before being given its own genus by naturalist Richard Lydekker in 1889. While many species have been assigned to ''Peloneustes'', ''P. philarchus'' is currently the only one still considered valid, with the others moved to different genera, considered '' nomina dubia'', or synonymised with ''P. philarchus''. Some of the material formerly assigned to ''P. evansi'' have since been reassigned to ''"Pliosaurus" andrewsi''. ''Peloneustes'' is known from many specimens, including some very complete material. With a total length of , ''Peloneustes'' is not a large pliosaurid. It had a large, triangular skull, which occupied about a fifth of its body length. T ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Rostrum (anatomy)
Rostrum (from Latin ', meaning ''beak'') is a term used in anatomy for a number of phylogenetically unrelated structures in different groups of animals. Invertebrates * In crustaceans, the rostrum is the forward extension of the carapace in front of the eyes. It is generally a rigid structure, but can be connected by a hinged joint, as seen in Leptostraca. * Among insects, the rostrum is the name for the piercing mouthparts of the order Hemiptera as well as those of the snow scorpionflies, among many others. The long snout of weevils is also called a rostrum. * Gastropod molluscs have a rostrum or proboscis. * Cephalopod molluscs have hard beak-like mouthparts referred to as the rostrum. File:Washington DC Zoo - Macrobrachium rosenbergii 6.jpg, Crustacean: the rostrum of the shrimp ''Macrobrachium rosenbergii'' is serrated along both edges. File:Gminatus australis with Beetle.jpg, Insect: assassin bug piercing its prey with its rostrum File:Architeuthis beak.jpg, Cephalopod: ...
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Strata
In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as either '' bedding surfaces'' or ''bedding planes''.Salvador, A. ed., 1994. ''International stratigraphic guide: a guide to stratigraphic classification, terminology, and procedure. 2nd ed.'' Boulder, Colorado, The Geological Society of America, Inc., 215 pp. . Prior to the publication of the International Stratigraphic Guide, older publications have defined a stratum as either being either equivalent to a single bed or composed of a number of beds; as a layer greater than 1 cm in thickness and constituting a part of a bed; or a general term that includes both ''bed'' and ''lamina''.Neuendorf, K.K.E., Mehl, Jr., J.P., and Jackson, J.A. , eds., 2005. ''Glossary of Geology'' 5th ed. Alexandria, Virginia, American Geological Institute. 779 pp. . ...
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Peloneustes Holotype Rostrum
''Peloneustes'' (meaning "mud swimmer") is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally described as a species of ''Plesiosaurus'' by palaeontologist Harry Govier Seeley in 1896, before being given its own genus by naturalist Richard Lydekker in 1889. While many species have been assigned to ''Peloneustes'', ''P. philarchus'' is currently the only one still considered valid, with the others moved to different genera, considered ''nomina dubia'', or synonymised with ''P. philarchus''. Some of the material formerly assigned to ''P. evansi'' have since been reassigned to ''"Pliosaurus" andrewsi''. ''Peloneustes'' is known from many specimens, including some very complete material. With a total length of , ''Peloneustes'' is not a large pliosaurid. It had a large, triangular skull, which occupied about a fifth of its body length. The ...
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Ichthyosaur
Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, although the term is now used more for the parent clade of the Ichthyosauria). Ichthyosaurs thrived during much of the Mesozoic era; based on fossil evidence, they first appeared around 250 million years ago ( Ma) and at least one species survived until about 90 million years ago, into the Late Cretaceous. During the Early Triassic epoch, ichthyosaurs and other ichthyosauromorphs evolved from a group of unidentified land reptiles that returned to the sea, in a development similar to how the mammalian land-dwelling ancestors of modern-day dolphins and whales returned to the sea millions of years later, which they gradually came to resemble in a case of convergent evolution. Ichthyosaurs were particularly abundant in the Late Triassic a ...
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Thalattosuchian
Thalattosuchia is a clade of marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Crocodilia and records from Thailand and China suggest that some members lived in freshwater. The clade contains two major subgroupings, the Teleosauroidea and Metriorhynchoidea. Teleosauroids are not greatly specialised for oceanic life, with back osteoderms similar to other crocodyliformes. Within Metriorhynchoidea, the Metriorhynchidae displayed extreme adaptions for life in the open ocean, including the transformation of limbs into flippers, the development of a tail fluke, and smooth, scaleless skin. Discovery and naming The term Thalattosuchia was coined by Fraas in 1901.Fraas E. 1901. Die Meerkrokodile (Thalattosuchia n. g.) eine neue Sauriergruppe der Juraformation. ''Jahreshefte des Vereins für vaterländische Naturkunde, Württemb ...
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Invertebrate
Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate subphylum Vertebrata. Familiar examples of invertebrates include arthropods, mollusks, annelids, echinoderms and cnidarians. The majority of animal species are invertebrates; one estimate puts the figure at 97%. Many invertebrate taxa have a greater number and variety of species than the entire subphylum of Vertebrata. Invertebrates vary widely in size, from 50  μm (0.002 in) rotifers to the 9–10 m (30–33 ft) colossal squid. Some so-called invertebrates, such as the Tunicata and Cephalochordata, are more closely related to vertebrates than to other invertebrates. This makes the invertebrates paraphyletic, so the term has little meaning in taxonomy. Etymology The word "invertebrate" comes from the Latin word ''vertebra'', whi ...
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Epicontinental Sea
An inland sea (also known as an epeiric sea or an epicontinental sea) is a continental body of water which is very large and is either completely surrounded by dry land or connected to an ocean by a river, strait, or "arm of the sea". An inland sea will generally have higher salinity than a freshwater lake, but usually lower salinity than the open ocean. Definition What constitutes an "inland sea" is complex and somewhat necessarily vague. The United States Hydrographic Office defined it as "a body of water nearly or completely surrounded by land, especially if very large or composed of salt water". Geologic engineers Heinrich Ries and Thomas L. Watson say an inland sea is merely a very large lake. Rydén, Migula, and Andersson and Deborah Sandler of the Environmental Law Institute add that an inland sea is "more or less" cut off from the ocean. It may be semi-enclosed, or connected to the ocean by a strait or "arm of the sea". An inland sea is distinguishable from a bay in t ...
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Fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of living fish species are ray-finned fish, belonging to the class Actinopterygii, with around 99% of those being teleosts. The earliest organisms that can be classified as fish were soft-bodied chordates that first appeared during the Cambrian period. Although they lacked a true spine, they possessed notochords which allowed them to be more agile than their invertebrate counterparts. Fish would continue to evolve through the Paleozoic era, diversifying into a wide variety of forms. Many fish of the Paleozoic developed external armor that protected them from predators. The first fish with jaws appeared in the Silurian period, after which many (such as sharks) became formidable marine predators rather than just the prey of arthropods. Mos ...
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Subaqueous Flight
Aquatic locomotion or swimming is biologically propelled motion through a liquid medium. The simplest propulsive systems are composed of cilia and flagella. Swimming has evolved a number of times in a range of organisms including arthropods, fish, molluscs, amphibians, reptiles, birds, and mammals. Evolution of swimming Swimming evolved a number of times in unrelated lineages. Supposed jellyfish fossils occur in the Ediacaran, but the first free-swimming animals appear in the Early to Middle Cambrian. These are mostly related to the arthropods, and include the Anomalocaridids, which swam by means of lateral lobes in a fashion reminiscent of today's cuttlefish. Cephalopods joined the ranks of the nekton in the late Cambrian, and chordates were probably swimming from the Early Cambrian. Many terrestrial animals retain some capacity to swim, however some have returned to the water and developed the capacities for aquatic locomotion. Most apes (including humans), however, lost th ...
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Thalassophonea
Thalassophonea is an extinct clade of pliosaurids from the Middle Jurassic to the early Late Cretaceous (Callovian to Turonian) of Australia, Europe, North America and South America. ''Thalassophonea'' was erected by Roger Benson and Patrick Druckenmiller in 2013. The name is derived from Greek ''thalassa'' (θάλασσα), "sea", and ''phoneus'' (φονεύς), "murderer". It is a stem-based taxon defined as "all taxa more closely related to ''Pliosaurus brachydeirus'' than to ''Marmornectes candrewi''". It includes the short necked and large headed taxa that typify the family. Classification The following cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... follows an analysis by Benson & Druckenmiller (2014). References Middle Jurassic first appearances C ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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