Pambdelurion
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Pambdelurion
''Pambdelurion'' is an extinct genus of panarthropod from the Cambrian aged Sirius Passet site in northern Greenland. Like the morphologically similar ''Kerygmachela'' from the same locality, ''Pambdelurion'' is thought to be closely related to arthropods, combining characteristics of " lobopodians" with those of primitive arthropods. Description ''Pambdelurion'' was large for a Cambrian animal, and is estimated to have reached a length of . ''Omnidens'', an organism from China that closely resembles ''Pambdelurion'' and may even be synonymous with it, reached even larger sizes, estimated to be based on the proportions of ''Pambdelurion''. The head of ''Pambdelurion'' bore a large pair of frontal appendages, homologous to the antennae of onychophorans and frontal appendages of radiodonts. These frontal appendages were weakly muscled and relatively soft, suggesting they may have served primarily as sensory organs, rather than for grasping prey. Between the appendages are a pair ...
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Lobopodia
The lobopodians, members of the informal group Lobopodia (from the Greek language, Greek, meaning "blunt feet"), or the formally erected phylum Lobopoda Cavalier-Smith (1998), are panarthropods with stubby legs called lobopods, a term which may also be used as a common name of this group as well. While the definition of lobopodians may differ between literatures, it usually refers to a group of soft-bodied, worm-like fossil panarthropods such as ''Aysheaia'' and ''Hallucigenia''. The oldest near-complete fossil lobopodians date to the Lower Cambrian; some are also known from Ordovician, Silurian and Carboniferous Lagerstätten. Some bear toughened claws, plates or spines, which are commonly preserved as small carbonaceous fossil, carbonaceous or Small Shelly Fossils, mineralized microfossils in Cambrian strata. The grouping is considered to be paraphyletic, as the three living panarthropod groups (Arthropoda, Tardigrada and Onychophora) are thought to have evolved from lobopodian ...
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Omnidens
''Omnidens amplus'', meaning "large all-tooth", is an extinct species of large Cambrian animal known only from a series of large mouth apparatus, originally mistaken as the mouthparts of anomalocaridids. When first named, it was interpreted as a giant priapulid, but is now considered a panarthropod. Its mouth apparatus closely resembles that of the smaller gilled lobopodian ''Pambdelurion'', indicating it is likely to have been a close relative of that species, with which it may be synonymous. With a maximum estimated body length of , ''Omnidens'' is suggested to have been the largest known free-living Cambrian organism. ''Omnidens'' fossils are found in the Maotianshan Shales. Description ''Omnidens'' is only known from mouthparts. The preserved mouthparts would have formed a short muscular, potentially protrusible pharynx surrounded by circles of spiny sclerites, which were reminiscent of the scalids of priapulids, kinorhynchs, and loriciferans. The inside of the pharynx wa ...
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Dinocarida
DinocarididaGreek, "Terrible crabs" – sometimes informally spelt Dinocarida, but the second 'id' is linguistically correct – see is a proposed fossil taxon of basal arthropods that flourished in the Cambrian period with occasional OrdovicianVan Roy, P.; Briggs, D. E. G. (2011). "A giant Ordovician anomalocaridid". Nature 473 (7348): 510–513. doi:10.1038/nature09920. edit and Devonian records. Characterized by a pair of frontal appendages and series of body flaps, the name of Dinocaridids comes from Greek, "deinos" and "caris" ("terrible crab"), referring to the suggested role of some of these members as the largest marine predators of their time. Dinocaridids are occasionally referred to as the 'AOPK group' by some literatures, as the group compose of Radiodonta (''Anomalocaris'' and relatives), Opabiniidae (''Opabinia'' and relatives), ''Pambdelurion'' and ''Kerygmachela.'' It is most likely paraphyletic, with ''Kerygmachela'' and ''Pambdelurion'' more basal than the clad ...
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Radiodonta
Radiodonta is an extinct Order (biology), order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the last two originally refer to the family Anomalocarididae, which previously included all species of this order but is now restricted to only a few species. Radiodonts are distinguished by their distinctive frontal appendages, which are morphologically diverse and used for a variety of functions. Radiodonts included the earliest large predators known, but they also included sediment sifters and filter feeders. Some of the most famous species of radiodonts are the Cambrian taxa ''Anomalocaris, Anomalocaris canadensis'', ''Hurdia, Hurdia victoria'', ''Peytoia nathorsti'', ''Titanokorys gainesii, Titanokorys gainessii, Cambroraster, Cambroraster falcatus'' and ''Amplectobelua, Amplectobelua symbrachiata'', the Ordovician ''Aegirocassis, Aegiroca ...
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Opabinia
''Opabinia regalis'' is an extinct, stem group arthropod found in the Middle Cambrian Burgess Shale Lagerstätte (505 million years ago) of British Columbia. ''Opabinia'' was a soft-bodied animal, measuring up to 7 cm in body length, and its segmented trunk had flaps along the sides and a fan-shaped tail. The head shows unusual features: five eyes, a mouth under the head and facing backwards, and a clawed proboscis that probably passed food to the mouth. ''Opabinia'' probably lived on the seafloor, using the proboscis to seek out small, soft food. Free abstract at Fewer than twenty good specimens have been described; 3 specimens of ''Opabinia'' are known from the Greater Phyllopod bed, where they constitute less than 0.1% of the community. When the first thorough examination of ''Opabinia'' in 1975 revealed its unusual features, it was thought to be unrelated to any known phylum, or perhaps a relative of arthropod and annelid ancestors. However, later studies since late 19 ...
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Arthropoda
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Sirius Passet
Sirius Passet is a Cambrian Lagerstätte in Peary Land, Greenland. The Sirius Passet Lagerstätte was named after the Sirius sledge patrol that operates in North Greenland. It comprises six places in Nansen Land, on the east shore of J.P. Koch Fjord in the far north of Greenland. It was discovered in 1984 by A. Higgins of the Geological Survey of Greenland. A preliminary account was published by Simon Conway Morris and others in 1987 and expeditions led by J. S. Peel and Conway Morris have returned to the site several times between 1989 and the present. A field collection of perhaps 10,000 fossil specimens has been amassed. It is a part of the Buen Formation. Age The fauna is inevitably compared to that of the Burgess Shale, although it is probably ten to fifteen million years older – vs. ) – and more closely contemporaneous with the fauna of the Maotianshan shales from Chengjiang, which are dated to . Preservation The preservation of the Sirius Passet is traditionall ...
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Arthropods
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Panarthropods
Panarthropoda is a proposed animal clade containing the extant phyla Arthropoda, Tardigrada (water bears) and Onychophora (velvet worms). Panarthropods also include extinct marine legged worms known as lobopodians ("Lobopodia"), a paraphyletic group where the last common ancestor and basal members (stem-group) of each extant panarthropod phylum are thought to have risen. However the term "Lobopodia" is sometimes expanded to include tardigrades and onychophorans as well. Common characteristics of the Panarthropoda include a segmented body, paired ladder-like, ventral nervous system, and the presence of paired appendages correlated with body segments. Taxonomy Not all studies support the monophyly of Panarthropoda, but most do, including neuroanatomical, phylogenomic and palaeontological studies. At least a close relationship between onychophorans and arthropods is widely agreed upon, but the position of tardigrades is more controversial. Some phylogenomic studies have ...
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Panarthropoda
Panarthropoda is a proposed animal clade containing the extant phyla Arthropoda, Tardigrada (water bears) and Onychophora (velvet worms). Panarthropods also include extinct marine legged worms known as lobopodians ("Lobopodia"), a paraphyletic group where the last common ancestor and basal members (stem-group) of each extant panarthropod phylum are thought to have risen. However the term "Lobopodia" is sometimes expanded to include tardigrades and onychophorans as well. Common characteristics of the Panarthropoda include a segmented body, paired ladder-like, ventral nervous system, and the presence of paired appendages correlated with body segments. Taxonomy Not all studies support the monophyly of Panarthropoda, but most do, including neuroanatomical, phylogenomic and palaeontological studies. At least a close relationship between onychophorans and arthropods is widely agreed upon, but the position of tardigrades is more controversial. Some phylogenomic studies have f ...
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Kerygmachela
''Kerygmachela kierkegaardi'' is a gilled lobopodian from the Cambrian Stage 3 aged Sirius Passet Lagerstätte in northern Greenland. Its anatomy strongly suggests that it, along with its relative ''Pambdelurion whittingtoni'', was a close relative of radiodont (anomalocaridids) and euarthropods. The generic name "Kerygmachela" derives from the Greek words ''Kerygma'' (proclamation) and ''Chela'' (claw), in reference to the flamboyant frontal appendages. The specific name, "kierkegaardi" honors Danish philosopher Søren Kierkegaard. Morphology The head of ''Kerygmachela'' possesses a pair of well-developed frontal appendages which correspond to those of other dinocaridids and siberiid lobopodians. Each of them terminates in a series of long spines. A pair of sessile, slit-like compound eyes is located slightly behind the base of these appendages. A small anterior-facing mouth is located below the head and bears a pair of stylet-like structures. The head also possesses a ...
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