Paleoptera
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Paleoptera
The name Palaeoptera (from Greek ( 'old') + ( 'wing')) has been traditionally applied to those ancestral groups of winged insects (most of them extinct) that lacked the ability to fold the wings back over the abdomen as characterizes the Neoptera. The Diaphanopterodea, which are palaeopteran insects, had independently and uniquely convergent evolution, evolved a different wing-folding mechanism. Both mayfly, mayflies and dragonfly, dragonflies lack any of the smell centers in their brain found in Neoptera. Disputed status The complexities of the wing-folding mechanism, as well as the mechanical operation of the wings in flight (insect flight, indirect flight muscles), are such that it clearly indicates the Neoptera are a monophyletic lineage. The problem is that the plesiomorphic absence of wing-folding does not necessarily mean the Palaeoptera form a natural group – they may simply be an assemblage containing all insects, closely related or not, that "are not Neoptera", ...
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Palaeoptera
The name Palaeoptera (from Greek ( 'old') + ( 'wing')) has been traditionally applied to those ancestral groups of winged insects (most of them extinct) that lacked the ability to fold the wings back over the abdomen as characterizes the Neoptera. The Diaphanopterodea, which are palaeopteran insects, had independently and uniquely convergent evolution, evolved a different wing-folding mechanism. Both mayfly, mayflies and dragonfly, dragonflies lack any of the smell centers in their brain found in Neoptera. Disputed status The complexities of the wing-folding mechanism, as well as the mechanical operation of the wings in flight (insect flight, indirect flight muscles), are such that it clearly indicates the Neoptera are a monophyletic lineage. The problem is that the plesiomorphic absence of wing-folding does not necessarily mean the Palaeoptera form a natural group – they may simply be an assemblage containing all insects, closely related or not, that "are not Neoptera", ...
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Insect Taxonomy
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; In: potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within. Nearly all insects hatch from eggs. ...
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Winged Insects
The Pterygota ( grc, πτερυγωτός, pterugōtós, winged) are a subclass of insects that includes the winged insects. It also includes insect orders that are secondarily wingless (that is, insect groups whose ancestors once had wings but that have lost them as a result of subsequent evolution). The pterygotan group comprises almost all insects. The insect orders not included are the Archaeognatha (jumping bristletails) and the Zygentoma ( silverfishes and firebrats), two primitively wingless insect orders. Also not included is Entognatha, which consist of three orders no longer considered to be insects: Protura, Collembola, and Diplura. Systematics Traditionally, this group was divided into the infraclasses Paleoptera and Neoptera. The former are nowadays strongly suspected of being paraphyletic, and better treatments (such as dividing or dissolving the group) are presently being discussed. In addition, it is not clear how exactly the neopterans are related among eac ...
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Pterygota
The Pterygota ( grc, πτερυγωτός, pterugōtós, winged) are a subclass of insects that includes the winged insects. It also includes insect orders that are secondarily wingless (that is, insect groups whose ancestors once had wings but that have lost them as a result of subsequent evolution). The pterygotan group comprises almost all insects. The insect orders not included are the Archaeognatha (jumping bristletails) and the Zygentoma (silverfishes and firebrats), two primitively wingless insect orders. Also not included is Entognatha, which consist of three orders no longer considered to be insects: Protura, Collembola, and Diplura. Systematics Traditionally, this group was divided into the infraclasses Paleoptera and Neoptera. The former are nowadays strongly suspected of being paraphyletic, and better treatments (such as dividing or dissolving the group) are presently being discussed. In addition, it is not clear how exactly the neopterans are related among each o ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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Monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have tak ...
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Tree Of Life Web Project
The Tree of Life Web Project is an Internet project providing information about the diversity and phylogeny of life on Earth. This collaborative peer reviewed project began in 1995, and is written by biologists from around the world. The site has not been updated since 2011, however the pages are still accessible. The pages are linked hierarchically, in the form of the branching evolutionary tree of life, organized cladistically. Each page contains information about one particular group of organisms and is organized according to a branched tree-like form, thus showing hypothetical relationships between different groups of organisms. In 2009 the project ran into funding problems from the University of Arizona. Pages and Treehouses submitted took a considerably longer time to be approved as they were being reviewed by a small group of volunteers, and apparently, around 2011, all activities ended. History The idea of this project started in the late 1980s. David Maddison was wor ...
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Archedictyon
Archedictyon (from Greek language, Greek ''Arche'' meaning first, original, ancient, primitive, or most basic and ''dictyo-'' meaning Net (device), net or netlike, plural "archedictya") is a name given to a Hypothesis, hypothetical scheme of wing vein, venation proposed for the common ancestor of all insect flight, winged insects. Description The nature of the archedictyon is an important consideration in the Taxonomy (biology), taxonomic classification of the large, primitive Palaeozoic insects known as Palaeodictyoptera and was used by Adolphe-Théodore Brongniart for this purpose as early as 1854. A National Museum of Natural History database for the study of ants, wasps, bees and termites describes the archedictyon as: ...the primitive original vein network characterizing the wings of many of the most ancient insect fossils (T-B, after Needham; Mackerras, in Commonwealth Scientific and Industrial Research Organisation, CSIRO); in Mastotermitidae and Hodotermitidae (Isoptera), t ...
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Paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Damselflies
Damselflies are flying insects of the suborder Zygoptera in the order Odonata. They are similar to dragonflies, which constitute the other odonatan suborder, Anisoptera, but are smaller and have slimmer bodies. Most species fold the wings along the body when at rest, unlike dragonflies which hold the wings flat and away from the body. An ancient group, damselflies have existed since at least the Lower Permian, and are found on every continent except Antarctica. All damselflies are predatory insects; both nymphs and adults actively hunt and eat other insects. The nymphs are aquatic, with different species living in a variety of freshwater habitats including acidic bogs, ponds, lakes and rivers. The nymphs moult repeatedly, at the last moult climbing out of the water to undergo metamorphosis. The skin splits down the back, they emerge and inflate their wings and abdomen to gain their adult form. Their presence on a body of water indicates that it is relatively unpolluted, but the ...
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Dragonflies
A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threatens dragonfly populations around the world. Adult dragonflies are characterized by a pair of large, multifaceted compound eyes, two pairs of strong, transparent wings, sometimes with coloured patches, and an elongated body. Many dragonflies have brilliant iridescent or metallic colours produced by structural colouration, making them conspicuous in flight. An adult dragonfly's compound eyes have nearly 24,000 ommatidia each. Dragonflies can be mistaken for the closely related damselflies, which make up the other odonatan infraorder ( Zygoptera) and are similar in body plan though usually lighter in build; however, the wings of most dragonflies are held flat and away from the body, while damselflies hold their wings folded at rest, along ...
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Odonata
Odonata is an order of flying insects that includes the dragonflies and damselflies. Members of the group first appeared during the Triassic, though members of their total group, Odonatoptera, first appeared in Late Carboniferous. The two common groups are distinguished with dragonflies, placed in the suborder Epiprocta, usually being larger, with eyes together and wings up or out at rest, while damselflies, suborder Zygoptera, are usually smaller with eyes placed apart and wings along body at rest. All Odonata have aquatic larvae called naiads (nymphs), and all of them, larvae and adults, are carnivorous. The adults can land, but rarely walk. Their legs are specialised for catching prey. They are almost entirely insectivorous. Etymology and terminology Fabricius coined the term ''Odonata'' in 1793 from the Ancient Greek ( Ionic form of ) 'tooth'. One hypothesis is that it was because their maxillae are notably toothed. Most insects also have toothed mandibles. The wo ...
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