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Paroedura Tanjaka
''Paroedura tanjaka'' is a species of lizard in the family Gekkonidae. It is endemic to northwestern Madagascar, and is currently known only to occur in Bemaraha and Namoroka. This nocturnal species is found in karstic dry forests at night, and is known to dwell in caves during the day. It is one of the largest species in the genus ''Paroedura'', reaching up to 18 cm in length. The species was first discovered in 1996 and officially named in 2000. Because its range is extremely limited and its habitat is declining, it is listed as Endangered by the IUCN Red List. While it is not the subject of any conservative measures, its known locations have been designated as national parks. Taxonomy The species ''Paroedura tanjaka'' was first erected in 2000 by Nussbaum and Raxworthy. The holotype ( UMMZ 224225) is a mature male collected from Bemaraha Nature Reserve in Madagascar on 18 March 1996. Several other individuals from the Bemaraha and Namoroka reserves were collected as pa ...
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Ronald Archie Nussbaum
Ronald Archie Nussbaum (born February 9, 1942) is an American herpetologist. He works with evolutionary biology and ecology of amphibians and reptiles, including systematics of caecilians and salamanders. He is a professor emeritus at the University of Michigan. Education Nussbaum possesses a bachelor's, master's, and doctorate in biology from the University of Idaho, Central Washington University, and Oregon State University, respectively. Taxa described *''Amietophrynus'' Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green & Wheeler, 2006 *'' Amphiglossus anosyensis'' Raxworthy & Nussbaum, 1993 *'' Amphiglossus mandokava'' Raxworthy & Nussbaum, 1993 *'' Amphiglossus punctatus'' Raxworthy & Nussbaum, 1993 *''Atretochoana'' Nussbaum & Wilkinson, 1995 *''Boulengerula fischeri'' Nussbaum & Hinkel, 1994 *'' Brookesia ambreensis'' Raxworthy & Nussbaum, 1995 *''Brookesia antakarana'' ...
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Paroedura Manongavato
''Paroedura manongavato'' is a species of lizard in the family Gekkonidae. It was described as a distinct species in 2023, and while specimens have been known for years prior, they were formerly assigned to ''Paroedura bastardi''. It is a microendemic species found only in two locations (Anja Reserve and Tsaranoro) in Madagascar. Taxonomy ''Paroedura manongavato'' was formerly not recognized as a separate species from the Mocquard's Madagascar ground gecko (''Paroedura bastardi''), with collected specimens being referred to ''P. bastardi'' or ''P.'' sp. aff. ''bastardi'' in 2021. A study in 2023 found that ''P. manongavato'' is a distinct species using genetic and morphological data. The holotype specimen was collected on 27 November 2014 and represents an adult female. The specific name is derived from the Malagasy language, Malagasy words “''manonga''” (meaning “to climb”) and “''vato''” (meaning “rock”). Phylogenetic analysis has found this species to be a sis ...
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Karst Topography
Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant rocks, such as quartzite, given the right conditions. Subterranean drainage may limit surface water, with few to no rivers or lakes. However, in regions where the dissolved bedrock is covered (perhaps by debris) or confined by one or more superimposed non-soluble rock strata, distinctive karst features may occur only at subsurface levels and can be totally missing above ground. The study of ''paleokarst'' (buried karst in the stratigraphic column) is important in petroleum geology because as much as 50% of the world's hydrocarbon reserves are hosted in carbonate rock, and much of this is found in porous karst systems. Etymology The English word ''karst'' was borrowed from German in the late 19th century, which entered German much earlier. ...
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Nocturnal
Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed senses of hearing, smell, and specially adapted eyesight. Some animals, such as cats and ferrets, have eyes that can adapt to both low-level and bright day levels of illumination (see metaturnal). Others, such as bushbabies and (some) bats, can function only at night. Many nocturnal creatures including tarsiers and some owls have large eyes in comparison with their body size to compensate for the lower light levels at night. More specifically, they have been found to have a larger cornea relative to their eye size than diurnal creatures to increase their : in the low-light conditions. Nocturnality helps wasps, such as ''Apoica flavissima'', avoid hunting in intense sunlight. Diurnal animals, including squirrels and songbirds, are active du ...
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Congener (biology)
Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organisms or constituents of living organisms of being special or doing something special. Each animal or plant species is special. It differs in some way from all other species...biological specificity is the major problem about understanding life." Biological specificity within ''Homo sapiens'' ''Homo sapiens'' has many characteristics that show the biological specificity in the form of behavior and morphological traits. Morphologically, humans have an enlarged cranial capacity and more gracile features in comparison to other hominins. The reduction of dentition is a feature that allows for the advantage of adaptability in diet and survival. As a species, humans are culture dependent and much of human survival relies on the culture and so ...
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Canthus (herpetology)
In snakes and amphibians, the canthus, canthal ridge or ''canthus rostralis'',Spawls S, Branch B. 1995. ''The Dangerous Snakes of Africa''. Ralph Curtis Books. Dubai: Oriental Press. 192 pp. . is the angle between the flat crown of the head and the side of the head between the eye and the snout,Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers''. Krieger Publishing Company, Malabar, Florida. . or more specifically, between the supraocular scale and the rostral scale. It is defined as a sharp ridge in many viperids, but is rounded in most rattlesnakes Rattlesnakes are venomous snakes that form the genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting small anima ..., for example.Campbell JA, Lamar WW. 2004. The Venomous Reptiles of the Western Hemisphere. Comstock Publishing Associates, Ithaca and Lo ...
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Rostral Scale
The rostral scale, or rostral, in snakes and other scaled reptiles is the median plate on the tip of the snout that borders the mouth opening. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: Comstock Publishing Associates. (7th printing, 1985). 1,105 pp. (in two volumes). . It corresponds to the mental scale in the lower jaw. The term pertains to the rostrum, or nose. In snakes, the shape and size of this scale is one of many characteristics used to differentiate species from one another. Related scales *Nasorostral scale *Mental scale *Labial scales See also *Snake scales *Anatomical terms of location Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ... References {{Reflist Snake scales ...
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Snout–vent Length
Snout–vent length (SVL) is a morphometric measurement taken in herpetology from the tip of the snout to the most posterior opening of the cloacal slit (vent)."direct line distance from tip of snout to posterior margin of vent" It is the most common measurement taken in herpetology, being used for all amphibians, lepidosaurs, and crocodilians (for turtles, carapace length (CL) and plastral length (PL) are used instead). The SVL differs depending on whether the animal is struggling or relaxed (if alive), or various other factors if it is a preserved specimen. For fossils, an osteological correlate Osteological correlates are marks on the bones of animals that are made from the causal interactions of the soft-tissue Soft tissue is all the tissue in the body that is not hardened by the processes of ossification or calcification such as bo ... such as precaudal length must be used. When combined with weight and body condition, SVL can help deduce age and sex. Advantages Bec ...
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Extent Of Occurrence
Extent may refer to: Computing * Extent (file systems), a contiguous region of computer storage medium reserved for a file * Extent File System, a discontinued file system implementation named after the contiguous region * Extent, a chunk of storage space logical volume management uses internally to provide various device mappings * Extent, in computer programming, is the period during which a variable has a particular value Other * Extent, a technical description of the wingspan of a bird, bat, or other flying animal * Extent, a writ allowing a creditor to seize or assume temporary ownership of a debtor's property; also, the actual seizure in its execution * Map extent A map extent is the portion of area of a region shown in a map. The limits of a map extent are defined in the coordinate system of the map. In Western culture, map extents usually have a rectangular shape, so they are defined with a minimum and ma ..., the portion of a region shown in a map See also * Ext ...
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Dry Forests
The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive several hundred centimeters of rain per year, they have long dry seasons that last several months and vary with geographic location. These seasonal droughts have great impact on all living things in the forest. Deciduous trees predominate in most of these forests, and during the drought a leafless period occurs, which varies with species type. Because trees lose moisture through their leaves, the shedding of leaves allows trees such as teak and mountain ebony to conserve water during dry periods. The newly bare trees open up the canopy layer, enabling sunlight to reach ground level and facilitate the growth of thick underbrush. Trees on moister sites and those with access to ground water tend to be evergreen. Infertile sites also tend to ...
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Tsingy De Namoroka Strict Nature Reserve
Namoroka National Park, formerly known as Tsingy de Namoroka Strict Nature Reserve, is a national park located in the northwestern part of Madagascar in the Mahajanga Province, specifically, the Soalala District. History Namoroka Strict Nature Reserve was established in 1927The Tsingy de Namoroka National Park Sitesheet
. World Database on Protected Areas.
and became a special reserve in 1966. It forms a complex with the neighboring .


Climate and geography

Located in the northwestern part of Madagascar about south of Soalala, N ...
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Endemic
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, the Cape sugarbird is found exclusively in southwestern South Africa and is therefore said to be ''endemic'' to that particular part of the world. An endemic species can be also be referred to as an ''endemism'' or in scientific literature as an ''endemite''. For example '' Cytisus aeolicus'' is an endemite of the Italian flora. '' Adzharia renschi'' was once believed to be an endemite of the Caucasus, but it was later discovered to be a non-indigenous species from South America belonging to a different genus. The extreme opposite of an endemic species is one with a cosmopolitan distribution, having a global or widespread range. A rare alternative term for a species that is endemic is "precinctive", which applies to ...
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