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Pareiasaurid
Pareiasaurs (meaning "cheek lizards") are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with scutes which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct at the end of the Permian during the Permian-Triassic extinction event. Description Pareiasaurs ranged in size from long, and may have weighed up to . They were stocky, with short tails, small heads, robust limbs, and broad feet. The cow-sized species ''Bunostegos'', which lived 260 million years ago, is the earliest known example of a tetrapod with a fully erect posture as its legs were positioned directly under its body. Pareiasaurs were protected by bony scutes called osteoderms that were set into the skin. Their heavy skulls were ...
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Deltavjatia
''Deltavjatia'' was a pareiasauromorph procolophonoid from the Tatarian stage of the Permian time period. It had a large body of about in length. ''Deltavjatia'' was an herbivore and lived in what is now Russia. The first specimen of ''Deltavjatia'' was a specimen of a skull and lower mandible (PIN 2212/1), found in the Urpalov Formation in Kotelnich, Vyatka River. Since then, numerous mostly complete skeletons have been found, many of them being so well preserved due to the silty, anaerobic environment of the Kotelnich deposits that fossilised white blood cells are able to be distinguished in them. Analyses of the bone histology of ''Deltavjatia'' show that they grew very rapidly during the early stages of their ontogeny but that their growth rate drastically slowed down once they reached approximately half of their full body size. ''Deltavjatia'' was placed as a subtaxon of Pareiasauridae by M.S.Y. Lee in 1997. See also *Pareiasaur *Procolophonia *Kotelnich *Vyatka (river) ...
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Scutosaurus
''Scutosaurus'' ("shield lizard") is an extinct genus of pareiasaur parareptiles. Its genus name refers to large plates of armor scattered across its body. It was a large anapsid reptile that, unlike most reptiles, held its legs underneath its body to support its great weight. Fossils have been found in the Sokolki Assemblage Zone of the Malokinelskaya Formation in European Russia, close to the Ural Mountains, dating to the late Permian (Lopingian) between 264 and 252 million years ago. Research history The first fossils were uncovered by Russian paleontologist Vladimir Prokhorovich Amalitskii while documenting plant and animal species in the Upper Permian sediments in the Northern Dvina River, Arkhangelsk District, Northern European Russia. Amalitskii had discovered the site in 1899, and he and his wife Anne Amalitskii continued to oversee excavation until 1914, recovering numerous nearly complete and articulated (in their natural position) skeletons belonging to a menage ...
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Anthodon (reptile)
''Anthodon'' (meaning "flower tooth") is an extinct genus of pareiasaurid parareptile from the Permian period of South Africa and Tanzania. History Richard Owen, who described ''Anthodon'', thought it was a dinosaur because dinosaurian skull material from the Early Cretaceous had become associated with the Permian material. The dinosaur material was later separated out by Robert Broom in 1912 and was renamed as the stegosaurid ''Paranthodon'' by Franz Nopcsa in 1929. Description This small form combines the primitive feature of interpterygoid fenestrae with an advanced feature of turtle-like armor. It was about in length, and weighed around . Small dermal ossicles covered the body, while the pattern of armor plates on the back reminiscent of a turtle shell. The tail was further shortened relative to less derived forms. Some other forms are characterized by having smooth skulls and armor on the dorsal midline. Skull The skull was small, and the cheekbones unornamented as in oth ...
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Elginia
''Elginia'' is an extinct genus of pareiasaurid known from the Late Permian of Scotland and China. It was named for the area around Elgin in Scotland, which has yielded many fossils referred to as the Elgin Reptiles. Discovery The type species of ''Elginia, Elginia mirabilis,'' was first described in 1893 by E.T. Newton, after fellow geologists John Horne and Archibald Geikie informed him of several unusual specimens stored at the Elgin Museum in Scotland. The specimens were collected several years earlier from the coarse sandstones of the nearby Cutties Hillock Quarry. The quarry’s sandstone, the Cutties Hillock Sandstone Formation, is often (but not always) considered equivalent to the otherwise fossil-poor Hopeman Sandstone Formation of broader Scottish geology. Tetrapod remains occupy a narrow section of the sandstone, lying above a pebbly layer. The sandstones of Cutties Hillock were deposited at the very end of the Permian Period, based on a dicynodont fauna simi ...
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Provelosaurus
''Provelosaurus'' is an extinct Pareiasaur genus of the Late Permian found on the road between Aceguá and Bagé in the Paleorrota, Rio Grande do Sul, Brazil. Found in the Rio do Rasto Formation, aged about 260 million years.http://www.ufrgs.br/geociencias/paleo/pareiassauro2.html Museu da UFRGS The holotype specimen found measures in length.Juan C. Cisneros, Paula Dentzien-Dias and Heitor Francischini. 2021. "The Brazilian Pareiasaur Revisited". ''Frontiers in Ecology and Evolution'' Classification Originally described as a South American representative of the genus ''Pareiasaurus'', it was assigned to a new genus ''Provelosaurus'' by Lee (1997), who noted it shows more affinities with the small, highly derived, South African dwarf pareiasaurs (called ''Pumiliopareiasauria Velosauria is a group of Pareiasauria, pareiasaur Sauropsida, reptiles that existed in the late Permian period. They ranged in size from the 50-centimeter-long ''Pumiliopareia'' to the 3-meter-long ''S ...
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Bunostegos
''Bunostegos'' ("knobbly kullroof") is an extinct genus of pareiasaur parareptile from the Late Permian of the Agadez Region in Niger. The type species, ''Bunostegos akokanensis'', was named from the Moradi Formation in 2003. It was a cow-sized animal with a distinctive skull that had large bony knobs, similar in form to those of other pareiasaurs but far larger. The species appears to have lived in a desert in the centre of the supercontinent of Pangaea. Analysis of the limb bones (including the scapulocoracoid, humerus, radius, ulna, pelvis, and femur) was published in 2015, and revealed that ''Bunostegos'' walked upright on four limbs, with the body held above ground. This new information directly suggests that it could be the first tetrapod with a fully erect gait. Description The animal has been described as about the size of a modern cow with a knobbly skull and bony plate armor on its back." Its teeth show it to have been a plant eater. It lived in an isolated desert ...
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Embrithosaurus
''Embrithosaurus'' was a pareiasaur from the Permian of South Africa. Description ''Embrithosaurus'' was in length and in weight. The skull is relatively deep and narrow. The body is lightly armoured with thin, smooth dermal scutes. Species * ''E. schwarzi'' (Watson, 1914). The type species. This is the most advanced species of this genus, as indicated by the teeth, which have nine cusps (in three groups of three). In cladistic analyses it is used as the monotypal species for the genus. * ''E. alexanderi'' (Haughton and Boonstra, 1929). This species was made the type for ''"Dolichopareia"''. As the name indicates, the skull is long and narrow. This would seem to indicate a different lifestyle or diet to other parieasaurs. More recently, it has been used as the monotypal species for the genus ''Nochelesaurus'' (it is not clear what the status of ''Embrithosaurus strubeni'' is, this may be a further transitional species). In cladistic analyses, this species is phylogenetically in ...
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Arganaceras
''Arganaceras'' ("Argana horn") is a medium-sized pareiasaur from the Late Permian Ikakern Formation of Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to .... It was about in length and had a horn-like structure on its snout. References Pareiasaurs Lopingian reptiles of Africa Fossils of Morocco Fossil taxa described in 2005 Prehistoric reptile genera {{permian-reptile-stub ...
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Pumiliopareia
''Pumiliopareia'' is an extinct genus of pareiasauridae, pareiasaurid parareptile from the Permian period of South Africa. It is known from a complete skeleton with osteoderms. Description ''Pumiliopareia'' was about 50 cm in length with a 12 cm skull. It is the smallest known member of the pareiasaurs, measuring only a fifth as long as some of its larger relatives. Like ''Anthodon (reptile), Anthodon'', its body was entirely covered with osteoderms. In analyses that support a pareiasaur origin of turtles, the sister taxon of the testudines. However it specifically shares with turtles a single trait only: Ribs greatly expanded anteroposteriorly (i.e. wide). Classification Originally included under the genus ''Nanoparia'', it was given its own name by Lee 1997 who found it did not form a clade with ''Nanoparia luckhoffi'', the type species of that genus, and preferred to have monophyletic genera. ''Nanoparia'' may still be a paraphyletic genus, which is allowed in Linan ...
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Nanopareia
''Nanoparia'' is an extinct genus of pareiasaur that lived in the Permian. Description It was about 60 cm in length, and weighed around 8 to 10 kilograms. Classification This is an unusual small, spiny specialised form. The skull is very similar to that of Pareiasaurus and Romer considered it a synonym of the latter. Orlov however (in Osnovy Paleontology, the monumental multi-volume Russian textbook of Paleontology) placed it in the Elginiinae. Kuhn (1969) however argues that while resembling Elginia in the ossifications at the rear of the skull, it differs completely in proportions and would not seem to be related. Lee (1997) considers it a basal member of the dwarf pareiasaurs. ''Nanoparia'' is believed to be among the pareiasaur genera most closely related to turtles Turtles are an order (biology), order of reptiles known as Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major gro ...
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Pangaea
Pangaea or Pangea () was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous approximately 335 million years ago, and began to break apart about 200 million years ago, at the end of the Triassic and beginning of the Jurassic. In contrast to the present Earth and its distribution of continental mass, Pangaea was centred on the equator and surrounded by the superocean Panthalassa and the Paleo-Tethys Ocean, Paleo-Tethys and subsequent Tethys Ocean, Tethys Oceans. Pangaea is the most recent supercontinent to have existed and the first to be reconstructed by geologists. Origin of the concept The name "Pangaea" is derived from Ancient Greek ''pan'' (, "all, entire, whole") and ''Gaia (mythology), Gaia'' or Gaea (, "Mother goddess, Mother Earth, land"). The concept that the continents once formed a contiguous land mass was hypothesised, with ...
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Therapsid
Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, as opposed to the sprawling posture of many reptiles and salamanders. Therapsids evolved from "pelycosaurs", specifically within the Sphenacodontia, more than 279.5 million years ago. They replaced the "pelycosaurs" as the dominant large land animals in the Middle Permian through to the Early Triassic. In the aftermath of the Permian–Triassic extinction event, therapsids declined in relative importance to the rapidly diversifying reptiles during the Middle Triassic. The therapsids include the cynodonts, the group that gave rise to mammals ( Mammaliaformes) in the Late Triassic, around 225 million years ago. Of the non-mammalian therapsids, only cynodonts survived beyond the end of the Triassic, with the only other ...
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