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Osphradium
The osphradium is a pigmented chemosensory epithelium patch in the mantle cavity present in six of the eight extant classes of molluscs (it is absent in the scaphopoda and monoplacophora; among cephalopoda, only the nautilus has what appears to be a set of osphradia), on or adjacent to the ctenidia (gills). The main function of this organ is disputed but it is believed to be used to test incoming water for silt and possible food particles or, in some species, for sensing the presence of light. It is a popular idea among malacologists that the presence of an osphradium should be a molluscan synapomorphy. However, an osphradium is absent in monoplacophorans and scaphopods. Moreover, the differences in enervation of these patches suggest that the osphradium (as a patch enervated from the ctenidial nerve) may be different from another organ sometimes called the posterior sensory organ (PSO) with separate enervation from the lateral nerve cords. Both types of sensory organs are foun ...
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Molluscs
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastropods ...
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Monoplacophora
Monoplacophora , meaning "bearing one plate", is a polyphyletic superclass of molluscs with a cap-like shell inhabiting deep sea environments . Extant representatives were not recognized as such until 1952; previously they were known only from the fossil record, and were thought to have become extinct 375 million years ago. Although the shell of many monoplacophorans is limpet-like in shape, they are not gastropods, nor do they have any close relation to gastropods. Definition Discussion about monoplacophorans is made difficult by the slippery definition of the taxon; some authors take it to refer to all non-gastropod mollusks with a single shell, or all single-shelled mollusks with serially repeated units; whereas other workers restrict the definition to cap-shaped forms, excluding spiral and other shapes of shell. The inclusion of the gastropod-like Bellerophontoidea within the group is also contentious. One attempt to resolve this confusion was to separate out the predo ...
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Cephalopoda
A cephalopod is any member of the molluscan class Cephalopoda ( Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by '' Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ide ...
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Nautilus
The nautilus (, ) is a pelagic marine mollusc of the cephalopod family Nautilidae. The nautilus is the sole extant family of the superfamily Nautilaceae and of its smaller but near equal suborder, Nautilina. It comprises six living species in two genera, the type of which is the genus '' Nautilus''. Though it more specifically refers to species ''Nautilus pompilius'', the name chambered nautilus is also used for any of the Nautilidae. All are protected under CITES Appendix II. Depending on species, adult shell diameter is between 4 and 10 inches. Nautilidae, both extant and extinct, are characterized by involute or more or less convolute shells that are generally smooth, with compressed or depressed whorl sections, straight to sinuous sutures, and a tubular, generally central siphuncle.Kümmel, B. 1964. Nautiloidae-Nautilida, in the Treatise on Invertebrate Paleontology, Geological Society of America and Univ of Kansas Press, Teichert and Moore eds. Having survived relativel ...
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Ctenidium (mollusc)
A ctenidium is a respiratory organ or gill which is found in many molluscs. This structure exists in bivalves, cephalopods, Polyplacophorans (chitons), and in aquatic gastropods such as freshwater snails and marine snails. Some aquatic gastropods possess one ctenidium known as monopectinate and others have a pair of ctenidia known as bipectinate. A ctenidium is shaped like a comb or a feather, with a central part from which many filaments or plate-like structures protrude, lined up in a row. It hangs into the mantle cavity and increases the area available for gas exchange.Respiratory system
The apple snail. Retrieved 2012-04-20. The word is Latinized but is derived from the ''ktenidion'' which means "li ...
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Silt
Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension with water. Silt usually has a floury feel when dry, and lacks plasticity when wet. Silt also can be felt by the tongue as granular when placed on the front teeth (even when mixed with clay particles). Silt is a common material, making up 45% of average modern mud. It is found in many river deltas and as wind-deposited accumulations, particularly in central Asia, north China, and North America. It is produced in both very hot climates (through such processes as collisions of quartz grains in dust storms) and very cold climates (through such processes as glacial grinding of quartz grains.) Loess is soil rich in silt which makes up some of the most fertile agricultural land on Earth. However, silt is very vulnerable to erosion, and it has poor mechanical properties, making constr ...
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Malacologist
Malacology is the branch of invertebrate zoology that deals with the study of the Mollusca (mollusks or molluscs), the second-largest phylum of animals in terms of described species after the arthropods. Mollusks include snails and slugs, clams, and cephalopods, along with numerous other kinds, many of which have shells. One division of malacology, conchology, is devoted to the study of mollusk shells. Malacology derives . Fields within malacological research include taxonomy, ecology and evolution. Applied malacology studies medical, veterinary, and agricultural applications; for example, mollusks as vectors of disease, as in schistosomiasis. Archaeology employs malacology to understand the evolution of the climate, the biota of the area, and the usage of the site. In 1681, Filippo Bonanni wrote the first book ever published that was solely about seashells, the shells of marine mollusks. The book was entitled: In 1868, the German Malacological Society was founded. Zoologica ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis ...
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Nature (journal)
''Nature'' is a British weekly scientific journal founded and based in London, England. As a multidisciplinary publication, ''Nature'' features peer-reviewed research from a variety of academic disciplines, mainly in science and technology. It has core editorial offices across the United States, continental Europe, and Asia under the international scientific publishing company Springer Nature. ''Nature'' was one of the world's most cited scientific journals by the Science Edition of the 2019 '' Journal Citation Reports'' (with an ascribed impact factor of 42.778), making it one of the world's most-read and most prestigious academic journals. , it claimed an online readership of about three million unique readers per month. Founded in autumn 1869, ''Nature'' was first circulated by Norman Lockyer and Alexander Macmillan as a public forum for scientific innovations. The mid-20th century facilitated an editorial expansion for the journal; ''Nature'' redoubled its efforts in ...
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Cephalopod Zootomy
A cephalopod is any member of the molluscan class Cephalopoda ( Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by '' Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ide ...
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