Opiliones Penis
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Opiliones Penis
The penis of the Opiliones (harvestmen) is an intromittent organ that is not present in other arachnids. It consists of a long shaft (the truncus) and a terminal capsule called a glans, containing a stylus and ejaculatory duct. It may have from one to three muscles, or none as in the specialized lineage Grassatores, where the penis is operated by haemolymph pressure. In some members of the Cyphophthalmi, the structure has been termed a "spermatopositor". This denomination refers to a superficial similarity ( analogy) with vertebrate penises and does not have any phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... relationship with it. References External linksOpiliones reproductive anatomy Dr. Jefferey Schultz, University of MarylandHarvestman Spider penis, ''Thelbu ...
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Opiliones
The Opiliones (formerly Phalangida) are an order of arachnids colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs. , over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014. Representatives of each extant suborder can be found on all continents except Antarctica. Well-preserved fossils have been found in the 400-million-year-old Rhynie cherts of Scotland, and 305-million-year-old rocks in France. These fossils look surprisingly modern, indicating that their basic body shape developed very early on, and, at least in some taxa, has changed little since that time. Their phylogenetic position within the Arachnida is disputed; their closest relatives may be the mites (Acari) or the Novogenuata (the Scorpiones, Pseudoscorpiones, and Solifugae). Althou ...
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Intromittent Organ
An intromittent organ is any external organ of a male organism that is specialized to deliver sperm during copulation. Intromittent organs are found most often in terrestrial species, as most non-mammalian aquatic species fertilize their eggs externally, although there are exceptions. For many species in the animal kingdom, the male intromittent organ is a hallmark characteristic of internal fertilization. Species with intromittent organs Invertebrates Molluscs Male cephalopods have a specialized arm, the hectocotylus, which is inserted into the female's mantle cavity to deliver a spermatophore during copulation. In some species, the hectocotylus breaks off inside the female's mantle cavity; in others, it can be used repeatedly to copulate with different females. Arachnids In spiders the intromittent organs are the male pedipalps, even though these are not primarily sexual organs, but serve as indirect mating organs; in the male the pedipalps have hollow, clubbed tips, oft ...
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Arachnid
Arachnida () is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons. Almost all adult arachnids have eight legs, although the front pair of legs in some species has converted to a sensory function, while in other species, different appendages can grow large enough to take on the appearance of extra pairs of legs. The term is derived from the Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, who was turned into a spider. Almost all extant arachnids are terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 100,000 named species, of which 47,000 are species of spiders. Morphology Almost all adult arachnids have eight legs, unlike adult inse ...
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Glans
The glans (, plural "glandes" ; from the Latin word for " acorn") is a vascular structure located at the tip of the penis in male mammals or a homologous genital structure of the clitoris in female mammals. Structure The exterior structure of the glans consists of mucous membrane, which is usually covered by foreskin or clitoral hood in naturally developed genitalia. This covering, called the prepuce, is normally retractable in adulthood. The glans naturally joins with the frenulum of the penis or clitoris, as well as the inner labia in women, and the foreskin in uncircumcised men. In non-technical or sexual discussions, often the word "clitoris" refers to the external glans alone, excluding the clitoral hood, frenulum, and internal body of the clitoris. Similarly, phrases "tip" or "head" of the penis refers to the glans alone. Sex differences in humans In males, the glans is known as the glans penis, while in females the glans is known as the clitoral glans. In females, ...
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Stylus
A stylus (plural styli or styluses) is a writing utensil or a small tool for some other form of marking or shaping, for example, in pottery. It can also be a computer accessory that is used to assist in navigating or providing more precision when using touchscreens. It usually refers to a narrow elongated staff, similar to a modern ballpoint pen. Many styluses are heavily curved to be held more easily. Another widely used writing tool is the stylus used by blind users in conjunction with the slate for punching out the dots in Braille. Etymology The English word ''stylus'' has two plurals: ''styli'' and ''styluses''. The original Latin word was spelled ; the spelling ''stylus'' arose from an erroneous connection with Greek (), 'pillar'.''Oxford Latin Dictionary'', s.v. "stilus" (2012). The Latin word had several meanings, including "a long, sharply pointed piece of metal; the stem of a plant; a pointed instrument for incising letters; the stylus (as used in literary composi ...
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Grassatores
The Grassatores are the most diverse infraorder of the Laniatores. It includes over 3,500 species distributed mainly in the tropicsKury, A.B.(2003). Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). '' Revista Ibérica de Aracnología'', vol. especial monográfico 1: 1-337. They are characterized by the male genitalia without musculature, operated by hydraulic pressure and by the double tarsal claws of posterior legs. Superfamilies * Assamioidea Sørensen, 1886 * Phalangodoidea Simon, 1879 * Samooidea Samooidea is a large superfamily in the Grassatores group of harvestmen. It includes around 380 species distributed throughout the tropics. They are characterized by the complex male genitalia, with eversible complementary sclerites. The Samooid ... Sørensen, 1886 * Zalmoxoidea Sørensen, 1886 * Gonyleptoidea Sundevall, 1833 References External linksClassification of OpilionesA synoptic taxonomic arrangement of the order Opiliones, down to f ...
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Haemolymph
Hemolymph, or haemolymph, is a fluid, analogous to the blood in vertebrates, that circulates in the interior of the arthropod (invertebrate) body, remaining in direct contact with the animal's tissues. It is composed of a fluid plasma in which hemolymph cells called hemocytes are suspended. In addition to hemocytes, the plasma also contains many chemicals. It is the major tissue type of the open circulatory system characteristic of arthropods (e.g. arachnids, crustaceans and insects). In addition, some non-arthropods such as molluscs possess a hemolymphatic circulatory system. Oxygen-transport systems were long thought unnecessary in insects, but ancestral and functional hemocyanin has been found in the hemolymph. Insect "blood" generally does not carry hemoglobin, although hemoglobin may be present in the tracheal system instead and play some role in respiration. Method of transport In the grasshopper, the closed portion of the system consists of tubular hearts and an aorta ...
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Cyphophthalmi
Cyphophthalmi is a suborder of harvestmen, colloquially known as mite harvestmen. Cyphophthalmi comprises 36 genera, and more than two hundred described species. The six families are currently grouped into three infraorders: the Boreophthalmi, Scopulophthalmi, and Sternophthalmi. Description Cyphophthalmi are smaller than the more familiar "daddy long-legs" harvestmen, with adults ranging from 1 to 7 mm in length. Moreover, their legs are relatively short compared to most other harvestmen, typically shorter than the body. Some superficially resemble mites, which is where they get their common name. Their coloration is almost always some shade of brown, with a heavily sclerotized body, and they are quite inconspicuous, residing in leaf litter or in caves. Many Cyphophthalmi are eyeless, and presumably rely on olfactory cues to find food and mates. Very little is known about their behavior, though they likely subsist mostly by scavenging and preying on minute arthropods. They ...
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Analogy
Analogy (from Greek ''analogia'', "proportion", from ''ana-'' "upon, according to" lso "against", "anew"+ ''logos'' "ratio" lso "word, speech, reckoning" is a cognitive process of transferring information or meaning from a particular subject (the analog, or source) to another (the target), or a linguistic expression corresponding to such a process. In a narrower sense, analogy is an inference or an argument from one particular to another particular, as opposed to deduction, induction, and abduction, in which at least one of the premises, or the conclusion, is general rather than particular in nature. The term analogy can also refer to the relation between the source and the target themselves, which is often (though not always) a similarity, as in the biological notion of analogy. Analogy plays a significant role in problem solving, as well as decision making, argumentation, perception, generalization, memory, creativity, invention, prediction, emotion, explanation, concep ...
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Vertebrate
Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with currently about 69,963 species described. Vertebrates comprise such groups as the following: * jawless fish, which include hagfish and lampreys * jawed vertebrates, which include: ** cartilaginous fish (sharks, rays, and ratfish) ** bony vertebrates, which include: *** ray-fins (the majority of living bony fish) *** lobe-fins, which include: **** coelacanths and lungfish **** tetrapods (limbed vertebrates) Extant vertebrates range in size from the frog species ''Paedophryne amauensis'', at as little as , to the blue whale, at up to . Vertebrates make up less than five percent of all described animal species; the rest are invertebrates, which lack vertebral columns. The vertebrates traditionally include the hagfish, which do no ...
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Penis
A penis (plural ''penises'' or ''penes'' () is the primary sexual organ that male animals use to inseminate females (or hermaphrodites) during copulation. Such organs occur in many animals, both vertebrate and invertebrate, but males do not bear a penis in every animal species, and in those species in which the male does bear a so-called penis, the penises in the various species are not necessarily homologous. The term ''penis'' applies to many intromittent organs, but not to all. As an example, the intromittent organ of most cephalopoda is the hectocotylus, a specialized arm, and male spiders use their pedipalps. Even within the Vertebrata there are morphological variants with specific terminology, such as Hemipenis, hemipenes. In most species of animals in which there is an organ that might reasonably be described as a penis, it has no major function other than intromission, or at least conveying the sperm to the female, but in the Eutheria, placental mammals the peni ...
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