Operculum (bryozoa)
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Operculum (bryozoa)
In the bryozoan order Cheilostomatida, the operculum is a calcareous or chitinous lid-like structure that protects the opening through which the polypide protrudes. Many species have modified the operculum in specialized zooids (avicularia) to form a range of mandibles (probably for defense) or hair-like setae (probably for cleaning, or in some unattached species, such as '' Selenaria'', for locomotion). The cyclostome family Eleidae also convergently evolved an opercular structure during the Early Cretaceous to Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ....Taylor, P.D. 1994. Systematics of the melicerititid cyclostome bryozoans; introduction and the genera ''Elea'', ''Semielea'' and ''Repromultelea''. Bulletin of the Natural history Museum, Geology Series 50:1 ...
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Bryozoa
Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869living species are known. At least two genera are solitary (''Aethozooides'' and ''Monobryozoon''); the rest are colonial. The terms Polyzoa and Bryozoa were introduced in 1830 and 1831, respectively. Soon after it was named, another group of animals was discovered whose filtering mechanism looked similar, so it was included in Bryozoa until 1869, when the two groups were no ...
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Cheilostomatida
Cheilostomatida, also called Cheilostomata, is an order of Bryozoa in the class Gymnolaemata. They are exclusively marine, colonial invertebrate animals. Cheilostome colonies are composed of calcium carbonate and grow on a variety of surfaces, including rocks, shells, seagrass and kelps. The colony shapes range from simple encrusting sheets to erect branching and even unattached forms. As in other bryozoan groups, each colony is composed of a few to thousands of individual polypides. Each individual has a U-shaped gut, and no respiratory, circulatory, or nerve system. Unique among bryozoans, cheilostome polypides are housed in a box-shaped zooids, which do not grow larger once the zooid is mature. The opening through which the polypide protrudes is protected by a calcareous or chitinous lidlike structure, an operculum. Cheilostomes possess avicularia, which have modified the operculum into a range of mandibles (possibly for defense) or hair-like setae (possibly for cleaning). ...
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Calcareous
Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an adjectival term applied to anatomical structures which are made primarily of calcium carbonate, in animals such as gastropods, i.e., snails, specifically about such structures as the operculum, the clausilium, and the love dart. The term also applies to the calcium carbonate tests of often more or less microscopic Foraminifera. Not all tests are calcareous; diatoms and radiolaria have siliceous tests. The molluscs are calcareous, as are calcareous sponges ( Porifera), that have spicules which are made of calcium carbonate. In botany ''Calcareous grassland'' is a form of grassland characteristic of soils containing much calcium carbonate from underlying chalk or limestone rock. In medicine The term is used in pathology, for example i ...
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Chitin
Chitin ( C8 H13 O5 N)n ( ) is a long-chain polymer of ''N''-acetylglucosamine, an amide derivative of glucose. Chitin is probably the second most abundant polysaccharide in nature (behind only cellulose); an estimated 1 billion tons of chitin are produced each year in the biosphere. It is a primary component of cell walls in fungi (especially basidiomycetes and filamentous fungi), the exoskeletons of arthropods such as crustaceans and insects, the radulae, cephalopod beaks and gladii of molluscs and in some nematodes and diatoms. It is also synthesised by at least some fish and lissamphibians. Commercially, chitin is extracted from the shells of crabs, shrimps, shellfishes and lobsters, which are major by-products of the seafood industry. The structure of chitin is comparable to cellulose, forming crystalline nanofibrils or whiskers. It is functionally comparable to the protein keratin. Chitin has proved useful for several medicinal, industrial and biotechnological purpos ...
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Polypide
The polypide in bryozoans encompasses most of the organs and tissues of each individual zooid. This includes the tentacles, tentacle sheath, U-shaped digestive tract, musculature and nerve cells. It is housed in the zooidal exoskeleton, which in cyclostomes Cyclostome is a biological term (from the Greek for "round mouth") used in a few different senses: * for the taxon Cyclostomi, which comprises the extant jawless fishes: the hagfish (Myxini) and the lampreys (Petromyzontidae). This was thought fo ... is tubular and in cheilostomes is box-shaped. See also Bryozoan Anatomy References Bryozoology Protostome anatomy {{bryozoan-stub ...
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Zooid
A zooid or zoöid is a single animal that is part of a colonial animal. This lifestyle has been adopted by animals from separate unrelated taxa. Zooids are multicellular; their structure is similar to that of other solitary animals. The zooids can either be directly connected by tissue (e.g. corals, Catenulida, Siphonophorae, Pyrosome or Ectoprocta) or share a common exoskeleton (e.g. Bryozoa or Pterobranchia). The colonial organism as a whole is called a ''zoon'' , plural ''zoa'' (from Ancient Greek meaning animal; plural , ). Zooids can exhibit polymorphism. For instance, extant bryozoans may have zooids adapted for different functions, such as feeding, anchoring the colony to the substratum and for brooding embryos. However, fossil bryozoans are only known by the colony structures that the zooids formed during life. There are correlations between the size of some zooids and temperature. Variations in zooid size within colonies of fossils can be used as an indicator of ...
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Mandible
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movable bone of the skull (discounting the ossicles of the middle ear). It is connected to the temporal bones by the temporomandibular joints. The bone is formed prenatal development, in the fetus from a fusion of the left and right mandibular prominences, and the point where these sides join, the mandibular symphysis, is still visible as a faint ridge in the midline. Like other symphyses in the body, this is a midline articulation where the bones are joined by fibrocartilage, but this articulation fuses together in early childhood.Illustrated Anatomy of the Head and Neck, Fehrenbach and Herring, Elsevier, 2012, p. 59 The word "mandible" derives from the Latin word ''mandibula'', "jawbone" (literally "one used for chewing"), from ''wikt:mandere ...
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Seta
In biology, setae (singular seta ; from the Latin word for "bristle") are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Annelid setae are stiff bristles present on the body. They help, for example, earthworms to attach to the surface and prevent backsliding during peristaltic motion. These hairs make it difficult to pull a worm straight from the ground. Setae in oligochaetes (a group including earthworms) are largely composed of chitin. They are classified according to the limb to which they are attached; for instance, notosetae are attached to notopodia; neurosetae to neuropodia. Crustaceans have mechano- and chemosensory setae. Setae are especially present on the mouthparts of crustaceans and can also be found on grooming limbs. In some cases, setae are modified into scale like structures. Setae on the legs of krill and other small crustaceans help them to gather phytoplankton. It captures them and allows th ...
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Cyclostomatida
Cyclostomatida, or cyclostomata (also known as cyclostomes), are an ancient order of stenolaemate bryozoans which first appeared in the Lower Ordovician. It consists of 7+ suborders, 59+ families, 373+ genera, and 666+ species. The cyclostome bryozoans were dominant in the Mesozoic; since that era, they have decreased. Currently, cyclostomes seldom constitute more than 20% of the species recorded in regional bryozoan faunas. Taxonomy Traditionally, cyclostomes have been divided into two groups according to the skeletal organization. In free-walled (or double-walled) cyclostomes, the exterior frontal walls of the zooids are uncalcified; autozooids have either a polygonal aperture bounded by vertical interior walls, or a subcircular aperture in species with kenozooids filling the spaces between the autozooids. By contrast, fixed-walled (or single-walled) cyclostomes have much of the exterior frontal wall calcified; autozooids normally have a subcircular aperture located at or ...
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Convergent Evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is homoplasy. The recurrent evolution of flight is a classic example, as flying insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas '' homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different traits. Convergent evolution is similar to parallel evo ...
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