Ossicaulis Lachnopus
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Ossicaulis Lachnopus
''Ossicaulis'' is a ditypic genus of mushrooms in the family Lyophyllaceae. Taxonomy The genus was circumscribed in 1985 to contain the species originally described as ''Agaricus lignatilis'' by Christian Hendrik Persoon in 1801. This was considered the sole species in the genus until 2007, when Marco Contu validly published a description of '' O. lachnopus'' from collections in Italy. Recent analysis (2013) confirms that '' O. lachnopus'' and '' O. lignatilis'' should be considered distinct species. Molecular genetics analysis suggests that ''Ossicaulis'' is most closely related to the genera '' Asterophora'', '' Hypsizygus'', '' Lyophyllum'', and ''Tricholomella''. Description ''Ossicaulis'' species have gills that are adnexed, adnate or somewhat decurrent and a stipe that is centrally or laterally attached to the cap. The hyphal system is monomitic (meaning only generative hyphae are present), the trama is regular, and there are clamp connections in the ...
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Fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''t ...
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Decurrent
''Decurrent'' (sometimes decurring) is a term used in botany and mycology to describe plant or fungal parts that extend downward. In botany, the term is most often applied to leaf blades that partly wrap or have wings around the stem or petiole and extend down along the stem. A "decurrent branching habit" is a plant form common for shrubs and most angiosperm trees, contrasted with the excurrent or "cone-shaped crown" common among many gymnosperms. The decurrent habit is characterized by having weak apical dominance that eventually produces a rounded or spreading tree crown. Examples of trees with decurrent habit are most hardwood trees: oak, hickory, maple, etc.Claud L. Brown, Robert G. McAlpine, Paul P. Kormanik, "Apical Dominance and Form in Woody Plants: A Reappraisal", ''American Journal of Botany'', Vol. 54, No. 2 (February 1967), pp. 153–162, In mycology, the term is most often applied to the hymenophore of a basidiocarp (such as the lamellae or "gills" of a mushroo ...
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Deciduous
In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, after flowering; and to the shedding of ripe fruit. The antonym of ''deciduous'' in the botanical sense is evergreen. Generally, the term "deciduous" means "the dropping of a part that is no longer needed or useful" and the "falling away after its purpose is finished". In plants, it is the result of natural processes. "Deciduous" has a similar meaning when referring to animal parts, such as deciduous antlers in deer, deciduous teeth (baby teeth) in some mammals (including humans); or decidua, the uterine lining that sheds off after birth. Botany In botany and horticulture, deciduous plants, including trees, shrubs and herbaceous perennials, are those that lose all of their leaves for part of the year. This process is called abscissio ...
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Micrometre
The micrometre ( international spelling as used by the International Bureau of Weights and Measures; SI symbol: μm) or micrometer (American spelling), also commonly known as a micron, is a unit of length in the International System of Units (SI) equalling (SI standard prefix "micro-" = ); that is, one millionth of a metre (or one thousandth of a millimetre, , or about ). The nearest smaller common SI unit is the nanometre, equivalent to one thousandth of a micrometre, one millionth of a millimetre or one billionth of a metre (). The micrometre is a common unit of measurement for wavelengths of infrared radiation as well as sizes of biological cells and bacteria, and for grading wool by the diameter of the fibres. The width of a single human hair ranges from approximately 20 to . The longest human chromosome, chromosome 1, is approximately in length. Examples Between 1 μm and 10 μm: * 1–10 μm – length of a typical bacterium * 3–8 μm – width of ...
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Hymenium
The hymenium is the tissue layer on the hymenophore of a fungal fruiting body where the cells develop into basidia or asci, which produce spores. In some species all of the cells of the hymenium develop into basidia or asci, while in others some cells develop into sterile cells called cystidia (basidiomycetes) or paraphyses (ascomycetes). Cystidia are often important for microscopic identification. The subhymenium consists of the supportive hyphae from which the cells of the hymenium grow, beneath which is the hymenophoral trama, the hyphae that make up the mass of the hymenophore. The position of the hymenium is traditionally the first characteristic used in the classification and identification of mushrooms. Below are some examples of the diverse types which exist among the macroscopic Basidiomycota and Ascomycota. * In agarics, the hymenium is on the vertical faces of the gills. * In boletes and polypores, it is in a spongy mass of downward-pointing tubes. * In puffballs, ...
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Cystidia
A cystidium (plural cystidia) is a relatively large cell found on the sporocarp of a basidiomycete (for example, on the surface of a mushroom gill), often between clusters of basidia. Since cystidia have highly varied and distinct shapes that are often unique to a particular species or genus, they are a useful micromorphological characteristic in the identification of basidiomycetes. In general, the adaptive significance of cystidia is not well understood. Classification of cystidia By position Cystidia may occur on the edge of a lamella (or analogous hymenophoral structure) (cheilocystidia), on the face of a lamella (pleurocystidia), on the surface of the cap (dermatocystidia or pileocystidia), on the margin of the cap (circumcystidia) or on the stipe (caulocystidia). Especially the pleurocystidia and cheilocystidia are important for identification within many genera. Sometimes the cheilocystidia give the gill edge a distinct colour which is visible to the naked eye or wit ...
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Cap Cuticle
The pileipellis is the uppermost layer of hyphae in the pileus of a fungal fruit body. It covers the trama, the fleshy tissue of the fruit body. The pileipellis is more or less synonymous with the cuticle, but the cuticle generally describes this layer as a macroscopic feature, while pileipellis refers to this structure as a microscopic layer. Pileipellis type is an important character in the identification of fungi. Pileipellis types include the cutis, trichoderm, epithelium, and hymeniderm types. Types Cutis A cutis is a type of pileipellis characterized by hyphae that are repent, that is, that run parallel to the pileus surface. In an ixocutis, the hyphae are gelatinous. Trichoderm In a trichoderm, the outermost hyphae emerge roughly parallel, like hairs, perpendicular to the cap surface. The prefix "tricho-" comes from a Greek word for "hair". In an ixotrichodermium, the outermost hyphae are gelatinous. Epithelium An epithelium is a pileipellis consisting of rounded ce ...
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Ellipsoidal
An ellipsoid is a surface that may be obtained from a sphere by deforming it by means of directional scalings, or more generally, of an affine transformation. An ellipsoid is a quadric surface;  that is, a surface that may be defined as the zero set of a polynomial of degree two in three variables. Among quadric surfaces, an ellipsoid is characterized by either of the two following properties. Every planar cross section is either an ellipse, or is empty, or is reduced to a single point (this explains the name, meaning "ellipse-like"). It is bounded, which means that it may be enclosed in a sufficiently large sphere. An ellipsoid has three pairwise perpendicular axes of symmetry which intersect at a center of symmetry, called the center of the ellipsoid. The line segments that are delimited on the axes of symmetry by the ellipsoid are called the ''principal axes'', or simply axes of the ellipsoid. If the three axes have different lengths, the figure is a triaxial ellipsoid (ra ...
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Basidiospore
A basidiospore is a reproductive spore produced by Basidiomycete fungi, a grouping that includes mushrooms, shelf fungi, rusts, and smuts. Basidiospores typically each contain one haploid nucleus that is the product of meiosis, and they are produced by specialized fungal cells called basidia. Typically, four basidiospores develop on appendages from each basidium, of which two are of one strain and the other two of its opposite strain. In gills under a cap of one common species, there exist millions of basidia. Some gilled mushrooms in the order Agaricales have the ability to release billions of spores. The puffball fungus ''Calvatia gigantea'' has been calculated to produce about five trillion basidiospores. Most basidiospores are forcibly discharged, and are thus considered ballistospores. These spores serve as the main air dispersal units for the fungi. The spores are released during periods of high humidity and generally have a night-time or pre-dawn peak concentration in the ...
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Clamp Connection
A clamp connection is a hook-like structure formed by growing hyphal cells of certain fungi. It is a characteristic feature of Basidiomycetes fungi. It is created to ensure that each cell, or segment of hypha separated by septa (cross walls), receives a set of differing nuclei, which are obtained through mating of hyphae of differing sexual types. It is used to maintain genetic variation within the hypha much like the mechanisms found in crozier (hook) during sexual reproduction. Formation Clamp connections are formed by the terminal hypha during elongation. Before the clamp connection is formed this terminal segment contains two nuclei. Once the terminal segment is long enough it begins to form the clamp connection. At the same time, each nucleus undergoes mitotic division to produce two daughter nuclei. As the clamp continues to develop it uptakes one of the daughter (green circle) nuclei and separates it from its sister nucleus. While this is occurring the remaining nuclei ...
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Trama (mycology)
In mycology, the term trama is used in two ways. In the broad sense, it is the inner, fleshy portion of a mushroom's basidiocarp, or fruit body. It is distinct from the outer layer of tissue, known as the pileipellis or cuticle, and from the spore-bearing tissue layer known as the hymenium. In essence, the trama is the tissue that is commonly referred to as the "flesh" of mushrooms and similar fungi.Largent D, Johnson D, Watling R. 1977. ''How to Identify Mushrooms to Genus III: Microscopic Features''. Arcata, CA: Mad River Press. . pp. 60–70. The second use is more specific, and refers to the "hymenophoral trama" that supports the hymenium. It is similarly interior, connective tissue, but it is more specifically the central layer of hyphae running from the underside of the mushroom cap to the lamella or gill, upon which the hymenium rests. Various types have been classified by their structure, including trametoid, cantharelloid, boletoid, and agaricoid, with agaricoid the ...
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Hypha
A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one or more cells surrounded by a tubular cell wall. In most fungi, hyphae are divided into cells by internal cross-walls called "septa" (singular septum). Septa are usually perforated by pores large enough for ribosomes, mitochondria, and sometimes nuclei to flow between cells. The major structural polymer in fungal cell walls is typically chitin, in contrast to plants and oomycetes that have cellulosic cell walls. Some fungi have aseptate hyphae, meaning their hyphae are not partitioned by septa. Hyphae have an average diameter of 4–6 µm. Growth Hyphae grow at their tips. During tip growth, cell walls are extended by the external assembly and polymerization of cell wall components, and the internal production of new cell membrane. The S ...
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