Notothenioidei
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Notothenioidei
Notothenioidei is one of 19 suborders of the order Perciformes. The group is found mainly in Antarctic and Subantarctic waters, with some species ranging north to southern Australia and southern South America. Notothenioids constitute approximately 90% of the fish biomass in the continental shelf waters surrounding Antarctica. Evolution and geographic distribution The Southern Ocean has supported fish habitats for 400 million years; however, modern notothenioids likely appeared sometime after the Eocene epoch. This period marked the cooling of the Southern Ocean, resulting in the stable, ice-cold conditions that have persisted to the present day. Another key factor in the evolution of notothenioids is the preponderance of the Antarctic Circumpolar Current (ACC), a large, slow-moving current that extends to the seafloor and precludes most migration to and from the Antarctic region. These unique environmental conditions in concert with the key evolutionary innovation of Antif ...
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Southern Ocean
The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the World Ocean, generally taken to be south of 60° S latitude and encircling Antarctica. With a size of , it is regarded as the second-smallest of the five principal oceanic divisions: smaller than the Pacific, Atlantic, and Indian oceans but larger than the Arctic Ocean. Over the past 30 years, the Southern Ocean has been subject to rapid climate change, which has led to changes in the marine ecosystem. By way of his voyages in the 1770s, James Cook proved that waters encompassed the southern latitudes of the globe. Since then, geographers have disagreed on the Southern Ocean's northern boundary or even existence, considering the waters as various parts of the Pacific, Atlantic, and Indian oceans, instead. However, according to Commodore John Leech of the International Hydrographic Organization (IHO), recent oceanographic research has discovered the importance of Southern ...
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Channichthyidae
The crocodile icefish or white-blooded fish comprise a family (Channichthyidae) of notothenioid fish found in the Southern Ocean around Antarctica. They are the only known vertebrates to lack hemoglobin in their blood as adults. Icefish populations are known to reside in the Atlantic and Indian sectors of the Southern Ocean, as well as the continental shelf waters surrounding Antarctica. Water temperatures in these regions remain relatively stable, generally ranging from . One icefish, ''Champsocephalus esox,'' is distributed north of the Antarctic Polar Frontal Zone. At least 16 species of crocodile icefish are currently recognized, although eight additional species have been proposed for the icefish genus ''Channichthys.'' In February 2021, scientists discovered and documented a breeding colony of ''Neopagetopsis ionah'' icefish estimated to have 60 million active nests across an area of approximately 92 square miles at the bottom of the Weddell Sea in Antarctica. The majority ...
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Perciform
Perciformes (), also called the Percomorpha or Acanthopteri, is an order (biology), order or superorder of Actinopterygii, ray-finned fish. If considered a single order, they are the most numerous order of vertebrates, containing about 41% of all bony fish. Perciformes means "perch-like". Perciformes is an Order (biology), Order within the Clade Percomorpha consisting of "perch-like" Percomorphans. This group comprises over 10,000 species found in almost all aquatic ecosystems. The order contains about 160 families, which is the most of any order within the vertebrates. It is also the most variably sized order of vertebrates, ranging from the ''Schindleria brevipinguis'' to the marlin in the genus ''Makaira''. They first appeared and diversified in the Late Cretaceous. Among the well-known members of this group are perch and darters (Percidae), sea bass and groupers (Serranidae). Characteristics The Fish anatomy#Fins, dorsal and Fish anatomy#Fins, anal fins are divided into ...
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Perciformes
Perciformes (), also called the Percomorpha or Acanthopteri, is an order or superorder of ray-finned fish. If considered a single order, they are the most numerous order of vertebrates, containing about 41% of all bony fish. Perciformes means "perch-like". Perciformes is an Order within the Clade Percomorpha consisting of "perch-like" Percomorphans. This group comprises over 10,000 species found in almost all aquatic ecosystems. The order contains about 160 families, which is the most of any order within the vertebrates. It is also the most variably sized order of vertebrates, ranging from the ''Schindleria brevipinguis'' to the marlin in the genus ''Makaira''. They first appeared and diversified in the Late Cretaceous. Among the well-known members of this group are perch and darters (Percidae), sea bass and groupers (Serranidae). Characteristics The dorsal and anal fins are divided into anterior spiny and posterior soft-rayed portions, which may be partially or compl ...
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Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest continent, being about 40% larger than Europe, and has an area of . Most of Antarctica is covered by the Antarctic ice sheet, with an average thickness of . Antarctica is, on average, the coldest, driest, and windiest of the continents, and it has the highest average elevation. It is mainly a polar desert, with annual precipitation of over along the coast and far less inland. About 70% of the world's freshwater reserves are frozen in Antarctica, which, if melted, would raise global sea levels by almost . Antarctica holds the record for the lowest measured temperature on Earth, . The coastal regions can reach temperatures over in summer. Native species of animals include mites, nematodes, penguins, seals and tardigrades. Where vegetation o ...
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Globin
The globins are a superfamily of heme-containing globular proteins, involved in binding and/or transporting oxygen. These proteins all incorporate the globin fold, a series of eight alpha helical segments. Two prominent members include myoglobin and hemoglobin. Both of these proteins reversibly bind oxygen via a heme prosthetic group. They are widely distributed in many organisms. Structure Globin superfamily members share a common three-dimensional fold. This 'globin fold' typically consists of eight alpha helices, although some proteins have additional helix extensions at their termini. Since the globin fold contains only helices, it is classified as an all-alpha protein fold. The globin fold is found in its namesake globin families as well as in phycocyanins. The globin fold was thus the first protein fold discovered (myoglobin was the first protein whose structure was solved). Helix packaging The eight helices of the globin fold core share significant nonlocal st ...
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Antifreeze Protein
Antifreeze proteins (AFPs) or ice structuring proteins refer to a class of polypeptides produced by certain animals, plants, fungi and bacteria that permit their survival in temperatures below the freezing point of water. AFPs bind to small ice crystals to inhibit the growth and recrystallization of ice that would otherwise be fatal. There is also increasing evidence that AFPs interact with mammalian cell membranes to protect them from cold damage. This work suggests the involvement of AFPs in cold acclimatization. Non-colligative properties Unlike the widely used automotive antifreeze, ethylene glycol, AFPs do not lower freezing point in proportion to concentration. Rather, they work in a noncolligative manner. This phenomenon allows them to act as an antifreeze at concentrations 1/300th to 1/500th of those of other dissolved solutes. Their low concentration minimizes their effect on osmotic pressure. The unusual properties of AFPs are attributed to their selective af ...
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Heat Shock Response
The heat shock response (HSR) is a cell stress response that increases the number of molecular chaperones to combat the negative effects on proteins caused by stressors such as increased temperatures, oxidative stress, and heavy metals. In a normal cell, proteostasis (protein homeostasis) must be maintained because proteins are the main functional units of the cell. Many proteins take on a defined configuration in a process known as protein folding in order to perform their biological functions. If these structures are altered, critical processes could be affected, leading to cell damage or death. The heat shock response can be employed under stress to induce the expression of heat shock proteins (HSP), many of which are molecular chaperones, that help prevent or reverse protein misfolding and provide an environment for proper folding. Protein folding is already challenging due to the crowded intracellular space where aberrant interactions can arise; it becomes more difficult when ...
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Membrane Lipid
Membrane lipids are a group of compounds (structurally similar to fats and oils) which form the double-layered surface of all cells (lipid bilayer). The three major classes of membrane lipids are phospholipids, glycolipids, and cholesterol. Lipids are amphiphilic: they have one end that is soluble in water ('polar') and an ending that is soluble in fat ('nonpolar'). By forming a double layer with the polar ends pointing outwards and the nonpolar ends pointing inwards membrane lipids can form a 'lipid bilayer' which keeps the watery interior of the cell separate from the watery exterior. The arrangements of lipids and various proteins, acting as receptors and channel pores in the membrane, control the entry and exit of other molecules and ions as part of the cell's metabolism. In order to perform physiological functions, membrane proteins are facilitated to rotate and diffuse laterally in two dimensional expanse of lipid bilayer by the presence of a shell of lipids closely attached ...
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Lateral Line
The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial cells, known as hair cells, which respond to displacement caused by motion and transduce these signals into electrical impulses via excitatory synapses. Lateral lines serve an important role in schooling behavior, predation, and orientation. Fish can use their lateral line system to follow the vortices produced by fleeing prey. Lateral lines are usually visible as faint lines of pores running lengthwise down each side, from the vicinity of the gill covers to the base of the tail. In some species, the receptive organs of the lateral line have been modified to function as electroreceptors, which are organs used to detect electrical impulses, and as such, these systems remain closely linked. Most amphibian larvae and some fully aquatic adult ...
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Adaptations
In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the population during that process. Thirdly, it is a phenotypic trait or adaptive trait, with a functional role in each individual organism, that is maintained and has evolved through natural selection. Historically, adaptation has been described from the time of the ancient Greek philosophers such as Empedocles and Aristotle. In 18th and 19th century natural theology, adaptation was taken as evidence for the existence of a deity. Charles Darwin proposed instead that it was explained by natural selection. Adaptation is related to biological fitness, which governs the rate of evolution as measured by change in allele frequencies. Often, two or more species co-adapt and co-evolve as they develop adaptations that interlock with those of the ot ...
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Biochemicals
Biochemistry or biological chemistry is the study of chemical processes within and relating to living organisms. A sub-discipline of both chemistry and biology, biochemistry may be divided into three fields: structural biology, enzymology and metabolism. Over the last decades of the 20th century, biochemistry has become successful at explaining living processes through these three disciplines. Almost all areas of the life sciences are being uncovered and developed through biochemical methodology and research. Voet (2005), p. 3. Biochemistry focuses on understanding the chemical basis which allows biological molecules to give rise to the processes that occur within living cells and between cells,Karp (2009), p. 2. in turn relating greatly to the understanding of tissues and organs, as well as organism structure and function.Miller (2012). p. 62. Biochemistry is closely related to molecular biology, which is the study of the molecular mechanisms of biological phenomena.Astb ...
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