Neoceratopsia
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Neoceratopsia
Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, ''Yinlong downsi'', lived between 161.2 and 155.7 million years ago.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> The last ceratopsian species, ''Triceratops prorsus'', became extinct during the Cretaceous–Paleogene extinction event, . ''Triceratops'' is by far the best-known ceratopsian to the general public. It is traditional for ceratopsian genus names to end in "''-ceratops''", although this is not always the case. One of the first named genera was ''Ceratops'' itself, which lent its name to the group, although it is considered a ''nomen dubium'' today as its fossil remains have no distinguishing characte ...
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Protoceratopsidae
Protoceratopsidae is a family of basal (primitive) ceratopsians from the Late Cretaceous period. Although ceratopsians have been found all over the world, protoceratopsids are only definitively known from Cretaceous strata in Asia, with most specimens found in China and Mongolia. As ceratopsians, protoceratopsids were herbivorous, with constantly replacing tooth batteries made for slicing through plants and a hooked beak for grabbing them. Protoceratopsids were small ceratopsians around 1-2.5 m in length. Their bony frill and horns were much smaller than more derived members of Ceratopsia, such as ceratopsids. Description Protoceratopsids were relatively small ceratopsians, averaging around 1-2.5 m in length from head to tail. Protoceratopsids have a frill and rostral bone characteristic of all ceratopsians. Their snout is particularly wedge-shaped with tall and narrow nostrils situated high on it. The antorbital fenestra is unusually small, and the antorbital fossa sits high on ...
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Archaeoceratops
''Archaeoceratops'', meaning "ancient horned face", is a genus of basal neoceratopsian dinosaur from the Early Cretaceous (Aptian stage) of north central China. It appears to have been bipedal and quite small, reaching in length and in body mass. It had a comparatively large head but no horns, possessing only a small bony frill projecting from the back of its head. Discovery and species Two specimens were found in the Middle Gray Unit of the Xinminbao Group, Gongpoquan Basin of the Mazong Shan area of Gansu Province, north central China. The type species, ''A. oshimai'', was named by Dong Zhiming and Azuma in 1997. It is the first basal neoceratopsian discovered in this area. The type specimen, IVPP V11114, consists of a partially complete skeleton including skull, caudal vertebrae, pelvis, and most of a hind foot. The second specimen (paratype), IVPP V11115, consists of an incomplete skeleton with a relatively well preserved caudal series, a partial hind limb, and a co ...
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Aquilops
''Aquilops'' is an early herbivorous ceratopsian dinosaur dating from the Early Cretaceous of North America, approximately 109 million to 104 million years ago. The type species is ''A. americanus''. Description The skull is 84.2 mm long. The holotype is possibly not from a full-grown individual. A comparison with related species indicates it might have been at 60% of its adult length. Wedel estimated the total body length of ''Aquilops'' at 60 cm and its weight at 1.5 kg. The authors established some unique traits. The rostral, the bone core of the snout beak, curves downwards and has an arched keel on its top with a bump on the front. In front of the tooth row the upper jaw rim is over its total length concave in side view. The skull opening, the antorbital fenestra, is twice as long as it is tall and has a pointed rear, below the eye socket. Discovery and naming In 1997, paleontologist Scott Madsen found the single fossil, a partial skull, in Carbon County ...
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Triceratops
''Triceratops'' ( ; ) is a genus of herbivore, herbivorous Chasmosaurinae, chasmosaurine Ceratopsidae, ceratopsid dinosaur that first appeared during the late Maastrichtian stage of the Late Cretaceous Period (geology), period, about 68 million years ago in what is now North America. It is one of the last-known non-avian dinosaur genera, and became extinct in the Cretaceous-Paleogene extinction event, Cretaceous–Paleogene extinction event 66 million years ago. The name ''Triceratops'', which literally means 'three-horned face', is derived from the Ancient Greek, Greek words () meaning 'three', () meaning 'horn', and () meaning 'face'. Bearing a large bony neck frill, frill, three horn (anatomy), horns on the skull, and a large four-legged body, exhibiting convergent evolution with rhinoceroses and bovines, ''Triceratops'' is one of the most recognizable of all dinosaurs and the most well-known ceratopsid. It was also one of the largest, up to long and in body m ...
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Helioceratops
''Helioceratops'' is a genus of herbivorous neoceratopsian dinosaur from the Middle Cretaceous of China. Discovery and naming In 2000 and 2002, at the Liufangzi site of China's eastern Jilin province, excavations took place during which the jaws were found of a ceratopsian new to science. The type species ''Helioceratops brachygnathus'' was named and described in 2009 by Jin Liyong, Chen Jun, Zan Shuqin and Pascal Godefroit. The generic name means "sun horned face" from the Greek ''helios'', "sun", ''keras'', "horn" and ''ops'', "face". The reference is that the Sun rises in the East and ceratopsians also "rose" in the East; i.e. they originated in the Orient. The name also refers to a close relationship with '' Auroraceratops'', the "dawn ceratopian". The specific name means "short jaw" from the Greek βραχύς, ''brachys'' and γνάθος, ''gnathos'', in reference to the distinguishing short lower jaw. ''Helioceratops'' was discovered in a layer of the Quantou Forma ...
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Liaoceratops
''Liaoceratops'', meaning "Liaoning horned face", is a ceratopsian dinosaur believed to be an early relative of the horned ceratopsids. It lived in the Early Cretaceous, 126 million years ago. It was discovered in China by a team of American and Chinese scientists. ''Liaoceratops'' was much smaller than its later relatives, but offers a glimpse into the early evolution of this group of dinosaurs. Discoveries and naming ''Liaoceratops'' was discovered in the famous Liaoning Province of China, where several fossils of feathered dinosaurs have also been collected. The type species ''Liaoceratops yanzigouensis'' was in 2002 named and described by Xu Xing, Peter Makovicky, Wang Xiaolin, Mark Norell and You Hailu. The generic name is derived from Liaoning and the Greek ''keras'', "horn" and ''ops'', "face". The specific name refers to the town Yanzigou. The holotype IVPP V12738 has been found in the Yixian Formation dating from the Barremian. These beds have also yielded fossil ...
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Graciliceratops
''Graciliceratops'' (meaning "slender horn") is a genus of Ceratopsia#Phylogeny, neoceratopsian dinosaurs that lived in Asia during the Late Cretaceous period. Discovery and naming The holotype, ZPAL MgD-I/156, was discovered at the Bayan Shireh Formation in Mongolia, coming from the Sheeregeen Gashoon locality near Sainshand. The discoveries were made during field exploration by the Polish-Mongolian Palaeontological Expedition, in 1971. Four years later, in 1975, the specimen was described by Teresa Maryańska and Halszka Osmólska and referred to the genus ''Microceratops''. However, Paul Sereno noted that the referral for this specimen was injustified and overall, the genus lacked diagnosis, therefore, ''Microceratops'' (now named ''Microceratus'') was considered a nomen dubium. The referred specimen was redescribed by him, creating a new genus and species: ''Graciliceratops mongoliensis''. The holotype is fragmentary, consisting of a very fragmented skull with mandibles; ver ...
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Ceratopsidae
Ceratopsidae (sometimes spelled Ceratopidae) is a family of ceratopsian dinosaurs including ''Triceratops'', ''Centrosaurus'', and ''Styracosaurus''. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are known from western North America, which formed the island continent of Laramidia during most of the Late Cretaceous. Ceratopsids are characterized by beaks, rows of shearing teeth in the back of the jaw, elaborate nasal horns, and a thin parietal-squamosal shelf that extends back and up into a frill. The group is divided into two subfamilies—Chasmosaurinae and Centrosaurinae. The chasmosaurines are generally characterized by long, triangular frills and well-developed brow horns. The centrosaurines had well-developed nasal horns or nasal bosses, shorter and more rectangular frills, and elaborate spines on the back of the frill. These horns and frills show remarkable variation and are the principal means by which the various species have b ...
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Leptoceratopsidae
Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, but they were more primitive and generally smaller. Phylogeny Leptoceratopsidae was originally named by Franz Nopcsa von Felső-Szilvás in 1923 as a subfamily Leptoceratopsinae, and its type species is ''Leptoceratops gracilis''. Mackovicky, in 2001, defined it as a stem-based taxon and a family consisting of ''Leptoceratops gracilis'' and all species closer to ''Leptoceratops'' than to ''Triceratops horridus''.Makovicky, P.J. 2001. A ''Montanoceratops cerorhynchus'' (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta, In: Tanke, D.H. & Carpenter, K. (Eds.). ''Mesozoic Vertebrate Life''. Bloomington: Indiana University Press. Pp. 243-262. The cladogram below follows the topologies from a 2015 analysis by Yiming ...
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Psittacosaurus
''Psittacosaurus'' ( ; "parrot lizard") is a genus of extinct ceratopsian dinosaur from the Early Cretaceous of what is now Asia, existing between 126 and 101 million years ago. It is notable for being the most species-rich non-avian dinosaur genus. Up to 12 species are known, from across China, Mongolia, Russia, and Thailand. The species of ''Psittacosaurus'' were obligate bipeds at adulthood, with a high skull and a robust beak. One individual was found preserved with long filaments on the tail, similar to those of ''Tianyulong''. ''Psittacosaurus'' probably had complex behaviours, based on the proportions and relative size of the brain. It may have been active for short periods of time during the day and night, and had well-developed senses of smell and vision. ''Psittacosaurus'' was one of the earliest ceratopsians, but closer to ''Triceratops'' than ''Yinlong''. Once in its own family, Psittacosauridae, with other genera like ''Hongshanosaurus'', it is now considered to be s ...
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Serendipaceratops
''Serendipaceratops'' (meaning "serendipitous horned face") is a genus of herbivorous ornithischian dinosaur, possibly an ankylosaur, from the early Cretaceous Period of Australia. The type species, ''S. arthurcclarkei'', was named in 2003. Discovery and species The first known bone from ''Serendipaceratops'', an ulna (lower arm bone), was in 1993 discovered in Australia near Kilcunda, on the south-east coast of Victoria. The find took place during the "Dinosaur Cove" project, uncovering remains at the basis of a cliff face at the shoreline; the fossil was excavated in the "Arch", a small area of sea-floor protected from the waves by a dam. In 2003, the type species ''Serendipaceratops arthurcclarkei'' was named and described by Tom Rich and Patricia Vickers-Rich.T. Rich and P. Vickers-Rich. 2003. Protoceratopsian? ulnae from Australia. ''Records of the Queen Victoria Museum Launceston'' 113:1-12 Initially, the discoverers had not considered the possibility that the fossil migh ...
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Auroraceratops
''Auroraceratops'', meaning "dawn horned face", is a genus of bipedal basal neoceratopsian dinosaur, from the Early Cretaceous (Aptian age) of north central China. The etymology of the generic name refers to its status as an early ceratopsian and also to Dawn Dodson, wife of Peter Dodson, one of the palaeontologists who described it. Discovery and species The first specimen was found in the Xinminpu Group, Gongpoquan Basin of the Mazong Shan area of Gansu Province, north central China. As of 2021, fossils from more than 80 more individuals have been found, including complete skeletons. The specific descriptor, "''rugosus''", Latin for "rough", refers to the various rugose areas on the surface of both skull and jaw, namely the wrinkled expansion of the lacrimal bone distinctive of this animal. ''Auroraceratops'' is the second basal neoceratopsian to be found in the Mazong Shan area, after ''Archaeoceratops''. The type specimen, IG-2004-VD-001, consists of a nearly complete ...
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