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Nematoda
The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broad range of environments. Less formally, they are categorized as Helminths, but are taxonomically classified along with arthropods, tardigrades and other moulting animals in the clade Ecdysozoa, and unlike flatworms, have tubular digestive systems with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum. Nematode species can be difficult to distinguish from one another. Consequently, estimates of the number of nematode species described to date vary by author and may change rapidly over time. A 2013 survey of animal biodiversity published in the mega jour ...
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Nematode Systematics
The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broad range of environments. Less formally, they are categorized as Helminths, but are taxonomically classified along with arthropods, tardigrades and other moulting animals in the clade Ecdysozoa, and unlike flatworms, have tubular digestive systems with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum. Nematode species can be difficult to distinguish from one another. Consequently, estimates of the number of nematode species described to date vary by author and may change rapidly over time. A 2013 survey of animal biodiversity published in the mega jou ...
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Chromadorea
The Chromadorea are a class of the roundworm phylum, Nematoda. They contain a single subclass (Chromadoria) and several orders. With such a redundant arrangement, the Chromadoria are liable to be divided if the orders are found to form several clades, or abandoned if they are found to constitute a single radiation. Formerly, they were treated as a subclass in the paraphyletic "Adenophorea" assemblage, which has been mostly abandoned by modern authors. It is also suspected that the Chromadorea may not be monophyletic as delimited here; at least the Monhysterida seem to be a distinct and far more ancient lineage than the rest. Members of this class' bodies usually have annules, their amphids elaborate and spiral, and they all have three esophageal glands. They usually live in marine sediments, although they can live elsewhere. They have a more sophisticated pharynx than most roundworms. Members of this class can be identified by the presence of eight conserved signature inde ...
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Enoplea
Enoplea (enopleans) is a class, which with the classes SecernenteaTree of Life Web Project (ToL) (2002)Nematoda Version of January 1, 2002. Retrieved November 2, 2008. and Chromadorea make up the phylum Nematoda in current taxonomy. The Enoplea are considered to be a more ancestral group than the Chromadorea, and researchers have referred to its members as the "ancestrally diverged nematodes", compared to the "more recently diverged nematodes" of Chromadorea. Description The Enoplea are distinguished from the Chromadorea by a number of characteristics. The enoplean esophagus is cylindrical or "bottle-shaped", compared to the bulbous chromadorean esophagus. Enopleans have pocket-like amphids, while chromadoreans have amphids shaped like slits, pores, coils, or spirals. An enoplean is smooth or marked with fine lines, while a chromadorean may have rings, projections, or setae. The enoplean excretory system is simple, sometimes made up of a single cell, while chromadoreans h ...
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Ecdysozoa
Ecdysozoa () is a group of protostome animals, including Arthropoda ( insects, chelicerata, crustaceans, and myriapods), Nematoda, and several smaller phyla. They were first defined by Aguinaldo ''et al.'' in 1997, based mainly on phylogenetic trees constructed using 18S ribosomal RNA genes. A large study in 2008 by Dunn ''et al.'' strongly supported the Ecdysozoa as a clade, that is, a group consisting of a common ancestor and all its descendants. The group is also supported by morphological characters, and includes all animals that grow by ecdysis, moulting their cuticle. The group was initially contested by a significant minority of biologists. Some argued for groupings based on more traditional taxonomic techniques, while others contested the interpretation of the molecular data. Etymology The name ''Ecdysozoa'' stems etymologically . Characteristics The most notable characteristic shared by ecdysozoans is a three-layered cuticle (four in Tardigrada) composed ...
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Secernentea
Secernentea was a class of nematodes in the Classical Phylogeny System (Chitwood, 1958) and is no longer in use. This morphological-based classification system has been replaced by the Modern Phylogeny system, where taxonomy assignment is based on small subunit ribosomal DNA (SSU rDNA). Characteristics of Secernentea are: * Amphid apertures are pore/slit-like * Derids are present in some; located near nerve ring * Phasmids are present; posterior * Excretory system is tubular * Cuticle is striated in two to four layers; lateral field is present * Three esophageal glands; esophageal structure varies * Males generally have one testis * Caudal alae are common * Sensory papillae are cephalic only; may be caudal papillae in males * Mostly terrestrial * Rarely found in fresh or marine water Systematics Subclasses and orders of Secernentea are: Tree of Life Web Project (ToL) (2002)Nematoda. Version of January 1, 2002. Retrieved November 2, 2008. * Subclass Rhabditia (paraphyletic?) ...
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Nematomorpha
Nematomorpha (sometimes called Gordiacea, and commonly known as horsehair worms, hairsnakes, or Gordian worms) are a phylum of parasitoid animals superficially similar to nematode worms in morphology, hence the name. Most species range in size from long, reaching 2 metres in extreme cases, and in diameter. Horsehair worms can be discovered in damp areas, such as watering troughs, swimming pools, streams, puddles, and cisterns. The adult worms are free-living, but the larvae are parasitic on arthropods, such as beetles, cockroaches, mantises, orthopterans, and crustaceans. About 351 freshwater species are known and a conservative estimate suggests that there may be about 2000 freshwater species worldwide. The name "Gordian" stems from the legendary Gordian knot. This relates to the fact that nematomorphs often coil themselves in tight balls that resemble knots. Description and biology Nematomorphs possess an external cuticle without cilia. Internally, they have only longitudina ...
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Caenorhabditis Elegans
''Caenorhabditis elegans'' () is a free-living transparent nematode about 1 mm in length that lives in temperate soil environments. It is the type species of its genus. The name is a blend of the Greek ''caeno-'' (recent), ''rhabditis'' (rod-like) and Latin ''elegans'' (elegant). In 1900, Maupas initially named it '' Rhabditides elegans.'' Osche placed it in the subgenus ''Caenorhabditis'' in 1952, and in 1955, Dougherty raised ''Caenorhabditis'' to the status of genus. ''C. elegans'' is an unsegmented pseudocoelomate and lacks respiratory or circulatory systems. Most of these nematodes are hermaphrodites and a few are males. Males have specialised tails for mating that include spicules. In 1963, Sydney Brenner proposed research into ''C. elegans,'' primarily in the area of neuronal development. In 1974, he began research into the molecular and developmental biology of ''C. elegans'', which has since been extensively used as a model organism. It was the first mult ...
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Helminths
Parasitic worms, also known as helminths, are large macroparasites; adults can generally be seen with the naked eye. Many are intestinal worms that are soil-transmitted and infect the gastrointestinal tract. Other parasitic worms such as schistosomes reside in blood vessels. Some parasitic worms, including leeches and monogeneans, are ectoparasites thus, they are not classified as helminths, which are endoparasites. Parasitic worms live in and feed in living hosts. They receive nourishment and protection while disrupting their hosts' ability to absorb nutrients. This can cause weakness and disease in the host, and poses a global health and economic problem. Parasitic worms cannot reproduce entirely within their host's body; they have a life cycle that includes some stages that need to take place outside of the host. Helminths are able to survive in their mammalian hosts for many years due to their ability to manipulate the host's immune response by secreting immunomodu ...
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Animalia
Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a bilaterally symmetric body plan. The Bilateria include the protostomes, containing animals such as nematodes, arthropods, flatworms, annelids and molluscs, and the deuterostomes, containing the echinoderms ...
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Phylum
In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained un ...
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Ecosystem
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the system through photosynthesis and is incorporated into plant tissue. By feeding on plants and on one another, animals play an important role in the movement of matter and energy through the system. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter, decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to a form that can be readily used by plants and microbes. Ecosystems are controlled by external and internal factors. External factors such as climate, parent material which forms the soil and topography, control the overall structure of an ecosystem but are not themselves influenced by the ecosys ...
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Oceanic Trench
Oceanic trenches are prominent long, narrow topographic depressions of the ocean floor. They are typically wide and below the level of the surrounding oceanic floor, but can be thousands of kilometers in length. There are about of oceanic trenches worldwide, mostly around the Pacific Ocean, but also in the eastern Indian Ocean and a few other locations. The greatest ocean depth measured is in the Challenger Deep of the Mariana Trench, at a depth of below sea level. Oceanic trenches are a feature of the Earth's distinctive plate tectonics. They mark the locations of convergent plate boundaries, along which lithospheric plates move towards each other at rates that vary from a few millimeters to over ten centimeters per year. Oceanic lithosphere moves into trenches at a global rate of about 3 km2/yr. A trench marks the position at which the flexed, subducting slab begins to descend beneath another lithospheric slab. Trenches are generally parallel to and about from ...
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