Najash Rionegrina
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Najash Rionegrina
''Najash'' is an extinct genus of basal snake from the Late Cretaceous Candeleros Formation of Patagonia. Like a number of other Cretaceous and living snakes it retained hindlimbs, but ''Najash'' is unusual in having well-developed legs that extend outside the rib cage, and a pelvis connected to the spine. Discovery and Description Fossils of ''Najash'' were found in the terrestrial Candeleros Formation, in Rio Negro Province, Argentina, and date to roughly 90 million years ago. The skull and spine of ''Najash'' show primitive features that resemble other Cretaceous snakes, such as ''Dinilysia patagonica'' and Madtsoiidae. Also, several characteristics of the neck and tail of ''Najash'' and '' Dinilysia patagonica'' show how the body plan of snakes evolved from a lizard-like ancestor. ''Najash'' had not lost its sacrum, the pelvic bone composed of several fused vertebrae, nor its pelvic girdle, which are absent in modern snakes, and in all other known fossil snake ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia an ...
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Snake
Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads (cranial kinesis). To accommodate their narrow bodies, snakes' paired organs (such as kidneys) appear one in front of the other instead of side by side, and most have only one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca. Lizards have evolved elongate bodies without limbs or with greatly reduced limbs about twenty-five times independently via convergent evolution, leading to many lineages of legless lizards. These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, although this rule is not universal (see Amphisbaen ...
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Cretaceous Snakes
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth by t ...
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Phylogenetics
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and do ...
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Eupodophis
''Eupodophis'' is an extinct genus of snake from the Late Cretaceous period. It has two small hind legs and is considered a transitional form between Cretaceous lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia altho ...s and limbless snakes. The feature, described as vestigial, was most likely useless to ''Eupodophis''. The type species ''Eupodophis descouensi'' was named in 2000 and resides now in the paleontology section of the Mim Museum in Beirut, Lebanon. The specific name is dedicated to the French naturalist Didier Descouens. The fossilized specimen from which the description of the type species was based was 85 cm (33.5 in) long and is approximately 92 million years old. It was found in Cenomanian-age limestone near the al-Nammoura village in Lebanon belonging to ...
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Pachyrhachis
''Pachyrhachis'' (from el, παχύς , 'thick' and el, ῥάχῐς , 'spine') is an extinct genus of snake with well developed hind legs known from fossils discovered in Ein Yabrud, near Ramallah, in the central West Bank. It is a relatively small snake, measuring more than long at maximum. ''Pachyrhachis'' appears to have been an ancient marine snake; the fossils occur in a marine limestone deposit, and the thickened bone of the ribs and vertebrae would have functioned as ballast to decrease the buoyancy of the animal, allowing it to dive beneath the ancient Cretaceous seas that it once inhabited. ''Pachyrhachis'' is one of three genera of Cenomanian snakes with hindlimbs. Although many modern pythons and boas still retain remnants of legs, in the form of small spurs, the tiny legs of Pachyrhachis included a hip, knee, and ankle joint. ''Pachyrhachis'' was originally described by Haas (1979, 1980) who noted it had a puzzling melange of snake and lizard features; its status ...
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Haasiophis
''Haasiophis'', consisting of the sole species ''Haasiophis terrasanctus'', is an extinct genus of snakes with hind limbs. It is one of three genera of Cenomanian snakes known to have possessed hindlimbs. Etymology The genus was named in honor of the late paleontologist Georg Haas, who first began work in the fossils of Ein Yabrud and started the description of the genus before he died, plus the Greek "ophis", for snake . The species names is from the Latin "terrasanctus" meaning "holy land". Specimen material ''H. terrasanctus'' is known from a single fossil discovered at Ein Yabrud in the Judean hills, near Ramallah, 20 km north of Jerusalem, in the central West Bank. This site also produced the type and only specimen for the sister genus ''Pachyrhachis''. et al. 2003 "The Anatomy and Relationships of Haasiophis terrasanctus, a Fossil Snake with Well-Developed Hind Limbs from the Mid-Cretaceous of the Middle East" ''Journal of Paleontology'' 77(3):536-558 This ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the ...
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Pelvic Girdle
The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The pelvic region of the trunk includes the bony pelvis, the pelvic cavity (the space enclosed by the bony pelvis), the pelvic floor, below the pelvic cavity, and the perineum, below the pelvic floor. The pelvic skeleton is formed in the area of the back, by the sacrum and the coccyx and anteriorly and to the left and right sides, by a pair of hip bones. The two hip bones connect the spine with the lower limbs. They are attached to the sacrum posteriorly, connected to each other anteriorly, and joined with the two femurs at the hip joints. The gap enclosed by the bony pelvis, called the pelvic cavity, is the section of the body underneath the abdomen and mainly consists of the reproductive organs (sex organs) and the rectum, while the pelvic ...
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Hip Bone
The hip bone (os coxae, innominate bone, pelvic bone or coxal bone) is a large flat bone, constricted in the center and expanded above and below. In some vertebrates (including humans before puberty) it is composed of three parts: the ilium, ischium, and the pubis. The two hip bones join at the pubic symphysis and together with the sacrum and coccyx (the pelvic part of the spine) comprise the skeletal component of the pelvis – the pelvic girdle which surrounds the pelvic cavity. They are connected to the sacrum, which is part of the axial skeleton, at the sacroiliac joint. Each hip bone is connected to the corresponding femur (thigh bone) (forming the primary connection between the bones of the lower limb and the axial skeleton) through the large ball and socket joint of the hip. Structure The hip bone is formed by three parts: the ilium, ischium, and pubis. At birth, these three components are separated by hyaline cartilage. They join each other in a Y-shaped porti ...
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Sacrum
The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part of the pelvic cavity, between the two wings of the pelvis. It forms joints with four other bones. The two projections at the sides of the sacrum are called the alae (wings), and articulate with the ilium at the L-shaped sacroiliac joints. The upper part of the sacrum connects with the last lumbar vertebra (L5), and its lower part with the coccyx (tailbone) via the sacral and coccygeal cornua. The sacrum has three different surfaces which are shaped to accommodate surrounding pelvic structures. Overall it is concave (curved upon itself). The base of the sacrum, the broadest and uppermost part, is tilted forward as the sacral promontory internally. The central part is curved outward toward the posterior, allowing greater room for the p ...
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Body Plan
A body plan, ( ), or ground plan is a set of morphological features common to many members of a phylum of animals. The vertebrates share one body plan, while invertebrates have many. This term, usually applied to animals, envisages a "blueprint" encompassing aspects such as symmetry, layers, segmentation, nerve, limb, and gut disposition. Evolutionary developmental biology seeks to explain the origins of diverse body plans. Body plans have historically been considered to have evolved in a flash in the Ediacaran biota; filling the Cambrian explosion with the results, and a more nuanced understanding of animal evolution suggests gradual development of body plans throughout the early Palaeozoic. Recent studies in animals and plants started to investigate whether evolutionary constraints on body plan structures can explain the presence of developmental constraints during embryogenesis such as the phenomenon referred to as phylotypic stage. History Among the pioneering zoologist ...
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