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Nacholapithecus
''Nacholapithecus kerioi'' was an ape that lived 14-15 million years ago during the Middle Miocene. Fossils have been found in the Nachola formation in northern Kenya. The only member of the genus ''Nacholapithecus'', it is thought to be a key genus in early hominid evolution. Similar in body plan to ''Proconsul'', it had a long vertebral column with six lumbar vertebrae, no tail, a narrow torso, large upper limbs with mobile shoulder joints, and long feet. Together with other Kenyapithecinae such as ''Equatorius'', ''Kenyapithecus'', and '' Griphopithecus'', ''Nacholapithecus'' displayed synapomorphies with ''Anoiapithecus''. Taxonomy Nacholapithecus was initially classified as belonging in ''Kenyapithecus'', then attributed to ''Equatorius'' (with ''Equatorius'' perhaps grouped into a sub-family ''Equatorinae'', instead of both species in ''Afropithecini''), finally recognised by Ishida ''et al.'' (1999) as a separate genus. Classified perhaps as a member of the family ''Pr ...
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Hominid Evolution
Human evolution is the evolutionary process within the history of primates that led to the emergence of ''Homo sapiens'' as a distinct species of the hominid family, which includes the great apes. This process involved the gradual development of traits such as human bipedalism and language, as well as interbreeding with other hominins, which indicate that human evolution was not linear but a web.Human Hybrids
(PDF). Michael F. Hammer. ''Scientific American'', May 2013.
The study of human evolution involves scientific disciplines, including

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Human Evolution
Human evolution is the evolutionary process within the history of primates that led to the emergence of ''Homo sapiens'' as a distinct species of the hominid family, which includes the great apes. This process involved the gradual development of traits such as human bipedalism and language, as well as interbreeding with other hominins, which indicate that human evolution was not linear but a web.Human Hybrids
(PDF). Michael F. Hammer. ''Scientific American'', May 2013.
The study of human evolution involves scientific disciplines, including
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Kyoto University Museum
The opened in Kyōto, Japan, in 2001. It exhibits materials from the collection of some 2,600,000 objects built up by Kyoto University since its foundation as Kyoto Imperial University in 1897. Arranged in accordance with three main themes - natural, cultural, and technological history - the collection includes artefacts excavated from the Yamashina-ku, Kyoto, Yamashina that have been Cultural Property (Japan), designated a National Treasure (Japan), National Treasure, several Important Cultural Properties of Japan, Important Cultural Properties, and materials from a number of excavations in China and Korea. The museum is part of the University Museum Association of Kyoto, a network of fourteen university museums in the city. See also * List of National Treasures of Japan (archaeological materials) * Kyoto National Museum * Ōtani University Museum * Ryūkoku Museum * Doshisha University Historical Museum * Bukkyō University Museum of Religious Culture * Kyoto Museum for World P ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Kenya
) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , official_languages = Constitution (2009) Art. 7 ational, official and other languages"(1) The national language of the Republic is Swahili. (2) The official languages of the Republic are Swahili and English. (3) The State shall–-–- (a) promote and protect the diversity of language of the people of Kenya; and (b) promote the development and use of indigenous languages, Kenyan Sign language, Braille and other communication formats and technologies accessible to persons with disabilities." , languages_type = National language , languages = Swahili , ethnic_groups = , ethnic_groups_year = 2019 census , religion = , religion_year = 2019 census , demonym = ...
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Proconsul (mammal)
''Proconsul'' is an extinct genus of primates that existed from 21 to 14 million years ago during the Miocene epoch. Fossil remains are present in Eastern Africa including Kenya and Uganda. Four species have been classified to date: ''P. africanus'', ''P. gitongai'', ''P. major'' and ''P. meswae''. The four species differ mainly in body size. Environmental reconstructions for the Early Miocene ''Proconsul'' sites are still tentative and range from forested environments to more open, arid grasslands. The gibbon and great apes, including humans, are held in evolutionary biology to share a common ancestral lineage, which may have included ''Proconsul''. Its name, meaning "before Consul" (Consul being a certain chimpanzee that, at the time of the genus's discovery, was on display in London), implies that it is ancestral to the chimpanzee. It might also be ancestral to the rest of the apes. Description The genus had a mixture of Old World monkey and ape characteristics, so its p ...
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Equatorius
''Equatorius'' is an extinct genus of kenyapithecine primate found in central Kenya at the Tugen Hills. Thirty-eight large teeth belonging to this middle Miocene hominid in addition to a mandibular and partially complete skeleton dated 15.58 Ma and 15.36 Ma. were later found. Analysis The anatomical structures in part was seen to be similar to ''Afropithecus'' and ''Proconsul A proconsul was an official of ancient Rome who acted on behalf of a consul. A proconsul was typically a former consul. The term is also used in recent history for officials with delegated authority. In the Roman Republic, military command, or ...''. Nevertheless, anatomy and morphology suggested the genus had an increased terrestrial habitat. Taxonomy , using their previous published study of ''K.africanus'', based the separate definition on comparisons of gnathic and dental anatomy. The classification's validity was subsequently challenged.See McCrossin & Benefit's comment in Notes Reference ...
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Kenyapithecus
''Kenyapithecus wickeri'' is a fossil ape discovered by Louis Leakey in 1961 at a site called Fort Ternan in Kenya. The upper jaw and teeth were dated to 14 million years ago. One theory states that ''Kenyapithecus'' may be the common ancestor of all the great apes. More recent investigations suggest ''Kenyapithecus'' is more primitive than that and is only slightly more modern than ''Proconsul'', which is considered to be an ape. Evidence suggests that ''Kenyapithecus wickeri'' was one of the species that started a radiation of apes out of Africa. Morphology Impressed by ''Kenyapithecus''s modern-looking teeth, Leakey declared ''Kenyapithecus'' to be "a very early ancestor of man himself." ''Kenyapithecus'' possessed craniodental adaptations for hard object feeding including thicker molar enamel, and a large mandible, large premolars and upper incisors that are similar to those seen in living pitheciine monkeys. ''Kenyapithecus'' also possessed macaque-like limbs adapted ...
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Griphopithecus
''Griphopithecus'' is a prehistoric ape from the Miocene of Turkey and Central Europe. Description ''Griphopithecus'' has been consistently grouped with stem hominoids. The material therefore indicates the range of hominoid locomotor anatomy in mid-Miocene Europe, rather than a specifically crown hominoid anatomy. Species *'' Griphopithecus alpani'' Tekkaya, 1974 *'' Griphopithecus suessi'' Abel, 1902 See also *''Graecopithecus'' *''Pierolapithecus'' *''Anoiapithecus'' *''Ouranopithecus'' *''Sivapithecus'' *''Lufengpithecus'' *''Gigantopithecus'' *''Khoratpithecus'' *''Chororapithecus ''Chororapithecus'' is an extinct great ape from the Afar region of Ethiopia roughly 8 million years ago during the Late Miocene, comprising one species, ''C. abyssinicus''. It is known from 9 isolated teeth discovered in a 2005–2007 survey of ...'' References External links Planet of the Apes (Scientific American/August, 2003) {{Taxonbar, from=Q2553860 Miocene primates of Europe Fos ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Anoiapithecus
''Anoiapithecus'' is an extinct ape genus thought to be closely related to ''Dryopithecus''. Both genera lived during the Miocene, approximately 12 million years ago. Fossil specimens named by Salvador Moyà-Solà are known from the deposits from Spain.. The discoverers described ''Anoiapithecus brevirostris'' as a hominoid (superfamily Hominoidea) in the dryopithecine tribe. They believe that it has more modern traits than the Kenyapithecines from which Kenya's '' Kenyapithecus wickeri'' brings fragmentary information. The African specimens are considered a sister taxon to the hominids, and 2 million years younger European specimens must be from the time after these two groups split. This means that hominids may have evolved in Europe. The name comes from the Anoia River region in Catalonia, where the fossil was found. It has been given the nickname ''Lluc'' (since it is a male individual). The name Lluc is the Catalan form of Luke, which in Latin suggests "light", as this d ...
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