Myrtales
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Myrtales
The Myrtales are an order of flowering plants placed as a sister to the eurosids II clade as of the publishing of the ''Eucalyptus grandis'' genome in June 2014. The APG III system of classification for angiosperms still places it within the eurosids. This finding is corroborated by the placement of the Myrtales in the Malvid clade by the One Thousand Plant Transcriptomes Initiative. The following families are included as of APGIII: * Alzateaceae S. A. Graham * Combretaceae R. Br. ( leadwood family) * Crypteroniaceae A. DC. * Lythraceae J. St.-Hil. ( loosestrife and pomegranate family) * Melastomataceae Juss. (including Memecylaceae DC.) * Myrtaceae Juss. (myrtle family; including Heteropyxidaceae Engl. & Gilg, Psiloxylaceae Croizat) * Onagraceae Juss. (evening primrose and Fuchsia family) * Penaeaceae Sweet ex Guill. (including Oliniaceae Arn., Rhynchocalycaceae L. A. S. Johnson & B. G. Briggs) * Vochysiaceae A. St.-Hil. The Cronquist system gives essentially the same compo ...
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APG III System
The APG III system of flowering plant classification is the third version of a modern, mostly molecular-based, system of plant taxonomy being developed by the Angiosperm Phylogeny Group (APG). Published in 2009, it was superseded in 2016 by a further revision, the APG IV system. Along with the publication outlining the new system, there were two accompanying publications in the same issue of the Botanical Journal of the Linnean Society: * The first, by Chase & Reveal, was a formal phylogenetic classification of all land plants (embryophytes), compatible with the APG III classification. As the APG have chosen to eschew ranks above order, this paper was meant to fit the system into the existing Linnaean hierarchy for those that prefer such a classification. The result was that all land plants were placed in the class Equisetopsida, which was then divided into 16 subclasses and a multitude of superorders. * The second, by Haston ''et al.'', was a linear sequence of families followi ...
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Alzateaceae
''Alzatea verticillata'' is a small flowering tree, native to the Neotropics. It inhabits moist submontane forests from Costa Rica and Panama in Central America south to Peru and Bolivia in tropical South America. It is the sole species of genus ''Alzatea'' and family Alzateaceae. Description ''Alzatea verticillata'' has opposite, obovate or elliptical leaves. Its flowers are actinomorphic and bisexual, and lack a corolla. The flowers and fruit are similar to the Myrtaceae but the ovary is superior. The fruit is a loculicidal capsule. The closest relatives of ''Alzatea'' are in the families Penaeaceae, Oliniaceae, Rhynchocalycaceae of southern Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area .... References * Schönenberger, Jürg and Conti, Elena. ''Molecular phylogeny ...
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Myrtaceae
Myrtaceae, the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All species are woody, contain essential oils, and have flower parts in multiples of four or five. The leaves are evergreen, alternate to mostly opposite, simple, and usually entire (i.e., without a toothed margin). The flowers have a base number of five petals, though in several genera, the petals are minute or absent. The stamens are usually very conspicuous, brightly coloured, and numerous. Evolutionary history Scientists hypothesize that the family Myrtaceae arose between 60 and 56 million years ago (Mya) during the Paleocene era. Pollen fossils have been sourced to the ancient supercontinent Gondwana. The breakup of Gondwana during the Cretaceous period (145 to 66 Mya) geographically isolated disjunct taxa and allowed for rapid speciation; i ...
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Eurosids II
The rosids are members of a large clade (monophyletic group) of flowering plants, containing about 70,000 species, more than a quarter of all angiosperms. The clade is divided into 16 to 20 orders, depending upon circumscription and classification. These orders, in turn, together comprise about 140 families. Fossil rosids are known from the Cretaceous period. Molecular clock estimates indicate that the rosids originated in the Aptian or Albian stages of the Cretaceous, between 125 and 99.6 million years ago. Today's forests are highly dominated by rosid species, which in turn helped with diversification in many other living lineages. Additionally, rosid herbs and shrubs are also a significant part of arctic/alpine, temperate floras, aquatics, desert plants, and parasites. Name The name is based upon the name "Rosidae", which had usually been understood to be a subclass. In 1967, Armen Takhtajan showed that the correct basis for the name "Rosidae" is a description of a group of ...
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Vochysiaceae
Vochysiaceae is a plant family belonging to the order of Myrtales. Description Trees or shrubs with opposite leaves; flowers are zygomorph 1-(3)-5 merous; ovary inferior or superior; one fertile stamen; fruits samara or capsules. Biogeography Six of the eight genera are native to the Neotropics. The genera '' Erismadelphus'' and '' Korupodendron'' are native to West and Central Africa. Evolutionary history The family likely originated in South America. ''Erismadelphus'' is thought to have diverged from '' Erisma'' approximately 30 million years ago, and traveled to Africa as the result of long-distance dispersal. Systematics Vochysiaceae are closest to Myrtaceae.Conti, E., A. Litt, P.G. Wilson, S.A Graham, B.G. Briggs, L.A.S. Johnson, K.J. Sytsma. 1997. Interfamiliar relationships in Myrtales: molecular phylogeny and patterns of morphological evolution. Systematic Botany 22: 629-647 Vochysiaceae consist of 7 genera with 217 species. The family is classified in two tribes ...
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Sonneratiaceae
Sonneratiaceae were a family of flowering plants placed in the order Myrtales by the Cronquist system. They consisted of two genera, ''Sonneratia'' and ''Duabanga''. These are now generally placed in their own monotypic subfamilies of the family Lythraceae, making Sonneratiaceae superfluous. The family is named for French naturalist Pierre Sonnerat Pierre Sonnerat (18 August 1748 – 31 March 1814) was a French naturalist, colonial administrator, writer and explorer. He described numerous species of plants and animals on his travels and is honoured in the genus ''Sonneratia'' and in other .... References * Myrtales Historically recognized angiosperm families {{Myrtales-stub ...
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Crypteroniaceae
The Crypteroniaceae are a family (biology), family of flowering plant, flowering trees and shrubs. The family includes 13 species in three genus, genera, native to Indomalaya. Genera There are three genera of Crypteroniaceae, all native to Asian tropical forests: * ''Axinandra'' Thwaites * ''Crypteronia'' Blume * ''Dactylocladus'' Oliv. The genus ''Axinandra'' includes four species, one in Sri Lanka (''A. zeylanica'') and the others in Borneo and the Malay Peninsula. ''Crypteronia'' includes seven species, ranging from eastern India through Southeast Asia and southern China to the Malay Peninsula, Indonesia, and New Guinea. ''Dactylocladus'' consists of a single species, native to the lowland peat swamp forests of Borneo. Phylogeny Morphological analyses, supported by recent chloroplast DNA analysis, indicates that Crypteroniaceae are most closely related to four small Myrtales, myrtalean families, Penaeaceae, Oliniaceae, and Rhynchocalycaceae of southern Africa, and Alzatea ...
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Combretaceae
The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family includes about 530 species of trees, shrubs, and lianas in ca 10 genera. The family includes the leadwood tree, ''Combretum imberbe''. Three genera, ''Conocarpus'', ''Laguncularia'', and ''Lumnitzera'', grow in mangrove habitats (mangals). The Combretaceae are widespread in the subtropics and tropics. Some members of this family produce useful construction timber, such as idigbo from ''Terminalia ivorensis''. The commonly cultivated ''Quisqualis indica'' is now placed in the genus ''Combretum''. Many plants in the Quisqualis species contain the Non-proteinogenic amino acid excitotoxin Quisqualic acid, a potent AMPA agonist.Excitotoxic cell death and delayed rescue in human neurons derived from NT2 cells, M Munir, L Lu and P Mcgonigl, Journal of Neuroscience, Vol 15, 7847–7860 White mangroves The family name comes from the type genus ''Combretum''; it also ...
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Penaeaceae
The Penaeaceae are a family of evergreen, leathery-leaved shrubs and small trees, native to South Africa. The family has 29 species in 9 genera.Stevens, P. F. (2001 onwards)Penaeaceae.Angiosperm Phylogeny Website.
Version 12, July 2012 nd more or less continuously updated since Accessed online: 5 June 2013. The family Penaeaceae was expanded under the of classification with the inclusion of the genera '''' (formerly in the monogeneric ...
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Memecylaceae
Memecylaceae A. P. de Candolle, DC. was a family (biology), family of flowering plants. The family included about 430 species of trees and shrubs in seven genus, genera. Memecylaceae are widespread in the tropics. The family has now been included within the Melastomataceae under the APG III system of classification,. When classified alone, this family included the following genera: *''Lijndenia'' *''Memecylon'' *''Mouriri'' *''Pternandra'' *''Spathandra'' *''Votomita'' *''Warneckea'' References

Myrtales Historically recognized angiosperm families {{rosid-tree-stub ...
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Lythraceae
Lythraceae is a family of flowering plants, including 32 genera, with about 620 species of herbs, shrubs, and trees. The larger genera include ''Cuphea'' (275 spp.), ''Lagerstroemia'' (56), ''Nesaea'' (50), ''Rotala'' (45), and ''Lythrum'' (35). It also includes the pomegranate (''Punica granatum'', formerly in Punicaceae) and the water caltrop (''Trapa natans'', formerly in Trapaceae). Lythraceae has a worldwide distribution, with most species in the tropics, but ranging into temperate climate regions as well. The family is named after the type genus, ''Lythrum'', the loosestrifes (e.g. ''Lythrum salicaria'' purple loosestrife) and also includes henna (''Lawsonia inermis''). It now includes the pomegranate, formerly classed in a separate family Punicaceae. The family also includes the widely cultivated crape myrtle trees. Botanically, the leaves are usually in pairs (opposite), and the flower petals emerge from the rim of the calyx tube. The petals often appear crumpled. Ch ...
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Angiosperms
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils are in the ...
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