Moschorhinidae
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Moschorhinidae
Akidnognathidae is an extinct family of therocephalian therapsids from the Late Permian and Early Triassic of South Africa, Russia and China. The family includes many large-bodied therocephalians that were probably carnivorous, including '' Moschorhinus'' and ''Olivierosuchus''. One akidnognathid, ''Euchambersia'', may even have been venomous. Akidnognathids have robust skulls with a pair of large caniniform teeth in their upper jaws. The family is morphologically intermediate between the more basal therocephalian group Scylacosauridae and the more derived group Baurioidea. Classification History The first family-level name used to classify an akidnognathid was Euchambersidae, erected by South African paleontologist Lieuwe Dirk Boonstra in 1934,Boonstra L.D. 1934. "A contribution to the morphology of the mammal-like reptiles of the suborder Therocephalia". ''Annals of the South African Museum'', 31: 215–267 in reference for the genus ''Euchambersia'', which is possibly one ...
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Euchambersia
''Euchambersia'' is an extinct genus of therocephalian therapsids that lived during the Late Permian in what is now South Africa and China. The genus contains two species. The type species ''E. mirabilis'' was named by paleontologist Robert Broom in 1931 from a skull missing the lower jaw. A second skull, belonging to a probably immature individual, was later described. In 2022, a second species, ''E. liuyudongi'', was named by Jun Liu and Fernando Abdala from a well-preserved skull. It is a member of the family Akidnognathidae, which historically has also been referred by as the synonymous Euchambersiidae (named after ''Euchambersia''). ''Euchambersia'' was a small and short-snouted therocephalian, possessing large canines as is typical of the group. However, it is notable among therocephalians for possessing ridges on its canines and a large indentation in the side of the skull. It has been proposed that these structures supported a venom delivery mechanism. If this statement ...
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Late Permian
Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, a concept in econometrics Music * Late (album), ''Late'' (album), a 2000 album by The 77s * Late!, a pseudonym used by Dave Grohl on his ''Pocketwatch (album), Pocketwatch'' album * Late (rapper), an underground rapper from Wolverhampton * Late (song), "Late" (song), a song by Blue Angel * "Late", a song by Kanye West from ''Late Registration'' Other * Late (Tonga), an uninhabited volcanic island southwest of Vavau in the kingdom of Tonga * Late (The Handmaid's Tale), "Late" (''The Handmaid's Tale''), a television episode * LaTe, Laivateollisuus, Oy Laivateollisuus Ab, a defunct shipbuilding company * Late may refer to a person who is Dead See also

* * * ''Lates'', a genus of fish in the lates perch family * Later (other) ...
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Tetrapod
Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct therapsids and all extant mammals). Tetrapods evolved from a clade of primitive semiaquatic animals known as the Tetrapodomorpha which, in turn, evolved from ancient lobe-finned fish (sarcopterygians) around 390 million years ago in the Middle Devonian period; their forms were transitional between lobe-finned fishes and true four-limbed tetrapods. Limbed vertebrates (tetrapods in the broad sense of the word) are first known from Middle Devonian trackways, and body fossils became common near the end of the Late Devonian but these were all aquatic. The first crown-tetrapods (last common ancestors of extant tetrapods capable of terrestrial locomotion) appeared by the very early Carboniferous, 350 million years ago. The specific aquatic ancestors ...
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Annatherapsidus
''Annatherapsidus'' is an extinct genus of therocephalians. It was a fairly small animal with a length of 91 cm and a 22 cm skull. See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera tha ... References Prehistoric synapsids of Europe Akidnognathids Fossil taxa described in 1961 Taxa named by Oskar Kuhn Therocephalia genera {{paleo-therapsid-stub ...
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Scylacosuchus
''Scylacosuchus'' is an extinct genus of therocephalian therapsids. It was a predatory eutherocephalian that lived in Lopingian epoch. Its fossils were found in Orenburg Oblast of Russia. The type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ... is ''Scylacosuchus orenburgensis''. References * Valentin P. Tverdokhlebov, Galina I. Tverdokhlebova, Alla V. Minikh, Mikhail V. Surkov, and Michael J. Benton, (2005) Upper Permian vertebrates and their sedimentological context in the South Urals, Russia, ''Earth-Science Reviews'' 69 27-77 55 Eutherocephalians Therocephalia genera Lopingian synapsids of Europe Prehistoric synapsids of Europe Fossil taxa described in 1968 Taxa named by Leonid Petrovich Tatarinov {{paleo-therapsid-stub ...
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Eutherocephalia
Eutherocephalia ("true beast head") is an extinct clade of advanced therocephalian therapsids. Eutherocephalians are distinguished from the lycosuchids and scylacosaurids, two early therocephalian families. While lycosuchids and scyalosaurids became extinct by the end of the Permian period, eutherocephalians survived the Permian–Triassic extinction event. The group eventually became extinct in the Middle Triassic. Characteristics The Eutherocephalians evolved several mammal-like traits through convergent evolution with Cynodontia. Among those traits were the loss of palatine teeth and the reduction of the parietal eye. The latter organ is instrumental in thermoregulation among lizards and snakes, indicating both eutherocephalians and cynodonts were evolving toward a more active, homeothermic lifestyle, though the eye never fully disappeared in the eutherocephalians. Classification The clade Eutherocephalia contains the majority of therocephalians, yet the phylogenetic rela ...
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Lycosuchus
''Lycosuchus'' ("wolf crocodile") is an extinct genus of carnivorous therocephalians which lived in the Middle Permian 265—260 Ma existing for approximately . As a member of the Lycosuchidae, the genus represents one of the earliest diverging therocephalians. The type species ''Lycosuchus vanderrieti'' is known from a handful of well-preserved specimens featuring the cranium and lower jaw; the holotype US D173 housed at the University of Stellenbosch, South Africa, is a near complete occluded skull. Specimen MB.R. 995, housed at the Museum für Naturkunde Berlin, Germany, consists of a near complete and isolated lower jaw, along with a partial snout and brain case. With the help of μCT data, Pusch et al (2020) described the endocranial anatomy of ''Lycosuchus vanderrieti''. It was a medium-sized predator, reaching 1.2 m (3.8 ft) in length with a skull 23 cm long., typical of early therocephalians. ''L. vanderrieti'' bore two functional canines in each maxilla, ...
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Procynosuchus
''Procynosuchus'' (Greek: "Before dog crocodile") is an extinct genus of cynodonts from the Late Permian. It is considered to be one of the earliest and most basal cynodonts. It was 60 cm (2 ft) long. Remains of ''Procynosuchus'' have been found in Russia, Germany, Zambia and South Africa.T. S. Kemp: ''The Origin and Evolution of Mammals'' Oxford University Press, 2005. Paleobiology As one of the earliest cynodonts, ''Procynosuchus'' has many primitive features, but it also has features that distinguish it from all other early therapsids. Some of these features were interpreted by Kemp (1980) as adaptations for a semi-aquatic lifestyle. For example, the wide zygapophyses of the vertebrae allow for a high degree of lateral flexibility, and ''Procynosuchus'' may have used anguilliform locomotion, or eel-like undulation, to swim through the water. The tail of ''Procynosuchus'' is also unusually long for a cynodont. The long haemal arches would have given the tail a lar ...
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Scylacosauria
Scylacosauria is a clade of therocephalian therapsids. It includes the basal family Scylacosauridae and the infraorder Eutherocephalia. Scylacosauridae and Eutherocephalia form this clade to the exclusion of Lycosuchidae, the most basal therocephalian family. Thus, Scylacosauria includes all therocephalians except lycosuchids. Below is a cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... showing the phylogenetic position of Sylacosauria: References Therocephalia Guadalupian first appearances Middle Triassic extinctions Tetrapod unranked clades {{paleo-Therapsid-stub ...
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Therocephalia
Therocephalia is an extinct suborder of eutheriodont therapsids (mammals and their close relatives) from the Permian and Triassic. The therocephalians ("beast-heads") are named after their large skulls, which, along with the structure of their teeth, suggest that they were carnivores. Like other non-mammalian synapsids, therocephalians were once described as "mammal-like reptiles". Therocephalia is the group most closely related to the cynodonts, which gave rise to the mammals. This relationship takes evidence in a variety of skeletal features. The fossils of therocephalians are numerous in the Karoo of South Africa, but have also been found in Russia, China, Tanzania, Zambia, and Antarctica. Early therocephalian fossils discovered in Middle Permian deposits of South Africa support a Gondwanan origin for the group, which seems to have spread quickly across Earth. Although almost every therocephalian lineage ended during the great Permian–Triassic extinction event, a few represe ...
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Akidnognathus
''Akidnognathus'' is an extinct genus of therocephalians. See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera tha ... References The main groups of non-mammalian synapsids at Mikko's Phylogeny Archive Akidnognathids Fossil taxa described in 1918 Taxa named by Sidney H. Haughton Therocephalia genera {{paleo-therapsid-stub ...
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