Microsporophyll
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Microsporophyll
A sporophyll is a leaf that bears sporangia. Both microphylls and megaphylls can be sporophylls. In heterosporous plants, sporophylls (whether they are microphylls or megaphylls) bear either megasporangia and thus are called megasporophylls, or microsporangia and are called microsporophylls. The overlap of the prefixes and roots makes these terms a particularly confusing subset of botanical nomenclature. Sporophylls vary greatly in appearance and structure, and may or may not look similar to sterile leaves. Plants that produce sporophylls include: ''Alaria esculenta'', a brown alga which shows sporophylls attached near the base of the alga.Dickson, Carola I. 1963. ''British Seaweeds''. The Kew Series. Lycophytes, where sporophylls may be aggregated into strobili (''Selaginella'' and some ''Lycopodium'' and related genera) or distributed singly among sterile leaves (''Huperzia''). Sporangia are borne in the axil or on the adaxial surface of the sporophyll. In heterosporous members ...
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Strobilus
A strobilus (plural: strobili) is a structure present on many land plant species consisting of sporangia-bearing structures densely aggregated along a stem. Strobili are often called cones, but some botanists restrict the use of the term cone to the woody seed strobili of conifers. Strobili are characterized by a central axis (anatomically a stem) surrounded by spirally arranged or decussate structures that may be modified leaves or modified stems. Leaves that bear sporangia are called ''sporophylls'', while sporangia-bearing stems are called '' sporangiophores''. Lycophytes Some members of both of the two modern classes of Lycopodiophyta (Lycopodiopsida and Isoetopsida) produce strobili. In all cases, the lateral organs of the strobilus are microphylls, bearing sporangia. In other lycophytes, ordinary foliage leaves can act as sporophylls, and there are no organized strobili. Diphasiastrum complanatum strobilus (01).jpeg, Strobili of ''Diphasiastrum'' Lycopodium saururus.JPG, St ...
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Sporangia
A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cycle. Sporangia can produce spores by mitosis, but in nearly all land plants and many fungi, sporangia are the site of meiosis and produce genetically distinct haploid spores. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains haploid nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and germinate into haploid hyphae. Although sexual reproduction in fungi varies between phyla, for some fungi the sporangium plays an indirect role in sexual reproducti ...
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Cycad
Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants of a species are either male or female. Cycads vary in size from having trunks only a few centimeters to several meters tall. They typically grow very slowly and live very long. Because of their superficial resemblance, they are sometimes mistaken for Arecaceae, palms or ferns, but they are not closely related to either group. Cycads are gymnosperms (naked-seeded), meaning their fertilization, unfertilized seeds are open to the air to be directly fertilized by pollination, as contrasted with angiosperms, which have enclosed seeds with more complex fertilization arrangements. Cycads have very specialized pollinators, usually a specific species of beetle. Both male and female cycads bear cones (strobilus, strobili), somewhat similar to conife ...
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Plant Morphology
Phytomorphology is the study of the morphology (biology), physical form and external structure of plants.Raven, P. H., R. F. Evert, & S. E. Eichhorn. ''Biology of Plants'', 7th ed., page 9. (New York: W. H. Freeman, 2005). . This is usually considered distinct from plant anatomy, which is the study of the internal Anatomy, structure of plants, especially at the microscopic level. Plant morphology is useful in the visual identification of plants. Recent studies in molecular biology started to investigate the molecular processes involved in determining the conservation and diversification of plant morphologies. In these studies transcriptome conservation patterns were found to mark crucial ontogenetic transitions during the plant life cycle which may result in evolutionary constraints limiting diversification. Scope Plant morphology "represents a study of the development, form, and structure of plants, and, by implication, an attempt to interpret these on the basis of similarit ...
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Gnetophyte
Gnetophyta () is a division of plants (alternatively considered the subclass Gnetidae or order Gnetales), grouped within the gymnosperms (which also includes conifers, cycads, and ginkgos), that consists of some 70 species across the three relict genera: ''Gnetum'' (family Gnetaceae), ''Welwitschia'' (family Welwitschiaceae), and ''Ephedra'' (family Ephedraceae). Fossilized pollen attributed to a close relative of ''Ephedra'' has been dated as far back as the Early Cretaceous. Though diverse in the Early Cretaceous, only three families, each containing a single genus, are still alive today. The primary difference between gnetophytes and other gymnosperms is the presence of vessel elements, a system of conduits that transport water within the plant, similar to those found in flowering plants. Because of this, gnetophytes were once thought to be the closest gymnosperm relatives to flowering plants, but more recent molecular studies have brought this hypothesis into question. Thoug ...
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Aerial Stem Modification
Aerial stem modifications are modifications to the aerial stems, vegetative buds and floral buds of plants growing in different conditions and which perform functions such as climbing, protection, support, synthesis of food, or vegetative propagation. Aerial stem structures that undergo modifications to perform these special functions include tendrils, thorns, hooks, phylloclade, tuberous stems and bulbils. The auxiliary or the terminal part of the modified structures show their stem nature. Tendrils Some weak stemmed plants produce wiry, coiled, sensitive and delicate organs for climbing. They are called tendrils. These may develop from either the axillary bud or the terminal bud of the stem. In ''Passiflora'', the tendrils develop from the axillary bud. In ''Cissus quadrangularis'' and in ''Vitis vinifera'' the terminal bud develops into tendrils. Thorns These are hard, woody, pointed structures meant for protection. They are provided with vascular tissue, which may ...
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Equisetum
''Equisetum'' (; horsetail, snake grass, puzzlegrass) is the only living genus in Equisetaceae, a family of ferns, which reproduce by spores rather than seeds. ''Equisetum'' is a "living fossil", the only living genus of the entire subclass Equisetidae, which for over 100 million years was much more diverse and dominated the understorey of late Paleozoic forests. Some equisetids were large trees reaching to tall. The genus ''Calamites'' of the family Calamitaceae, for example, is abundant in coal deposits from the Carboniferous period. The pattern of spacing of nodes in horsetails, wherein those toward the apex of the shoot are increasingly close together, is said to have inspired John Napier to invent logarithms. Modern horsetails first appeared during the Jurassic period. A superficially similar but entirely unrelated flowering plant genus, mare's tail (''Hippuris''), is occasionally referred to as "horsetail", and adding to confusion, the name "mare's tail" is sometimes ap ...
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Psilotum
''Psilotum'' is a genus of fern-like vascular plants. It is one of two genera in the family Psilotaceae commonly known as whisk ferns, the other being ''Tmesipteris''. Plants in these two genera were once thought to be descended from the earliest surviving vascular plants, but more recent phylogenies place them as basal ferns, as a sister group to Ophioglossales. They lack true roots and leaves are very reduced, the stems being the organs containing photosynthetic and conducting tissue. There are only two species in ''Psilotum'' and a hybrid between the two. They differ from those in ''Tmesipteris'' in having stems with many branches and a synangium with three lobes rather than two. Description and life cycle Whisk ferns in the genus ''Psilotum'' lack true roots but are anchored by creeping rhizomes. The stems have many branches with paired enations, which look like small leaves but have no vascular tissue. Above these enations there are synangia formed by the fusion of three sp ...
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Ginkgo
''Ginkgo'' is a genus of non-flowering seed plants. The scientific name is also used as the English name. The order to which it belongs, Ginkgoales, first appeared in the Permian, 270 million years ago, and is now the only living genus within the order. The rate of evolution within the genus has been slow, and almost all its species had become extinct by the end of the Pliocene. The sole surviving species, ''Ginkgo biloba'' is only found in the wild in China, but is cultivated around the world. The relationships between ginkgos and other groups of plants are not fully resolved. Prehistory The ginkgo (''Ginkgo biloba'') is a living fossil, with fossils similar to the modern plant dating back to the Permian, 270 million years ago. The closest living relatives of the clade are the cycads, which share with the extant ''G. biloba'' the characteristic of motile sperm. The ginkgo and cycad lineages are thought to have an extremely ancient divergence dating to the early Ca ...
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Fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Onoclea Sensibilis
''Onoclea sensibilis'', the sensitive fern, also known as the bead fern, is a coarse-textured, medium to large-sized deciduous perennial fern. The name comes from its sensitivity to frost, the fronds dying quickly when first touched by it. It is sometimes treated as the only species in ''Onoclea'', but some authors do not consider the genus monotypic. Description The sterile and fertile fronds of ''Onoclea sensibilis'' have independent stalks originating from the same rhizome, quite different from other ferns. The bright, yellow-green trophophylls (sterile fronds) are deeply pinnatifid and are typically borne at intervals along the creeping rhizome. The sterile fronds are deciduous with trophopods, swollen bases, that serve as over winter storage organs. The sterile fronds of ''O. var. sensibilis'' have a length of 1–1.3 m (3–4 ft) with 5–11 pinnae, leaf pairs, evenly spaced along the stipe. ''O. var. interrupta Maxim.'' fronds are shorter, 20–50 cm (8–20 in) long ...
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