Metaspriggina
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Metaspriggina
''Metaspriggina'' is a genus of chordate initially known from two specimens in the Middle Cambrian Burgess Shale and 44 specimens found in 2012 at the Marble Canyon bed in Kootenay National Park. Whilst named after the Ediacaran organism ''Spriggina'', later work has shown the two to be unrelated. ''Metaspriggina'' is considered to represent a primitive chordate, possibly transitional between cephalochordates and the earliest vertebrates, albeit this has been questioned because it seems to possess most of the characteristics attributed to craniates. It lacked fins and it had a weakly developed cranium, but it did possess two well-developed upward-facing eyes with nostrils behind them. ''Metaspriggina'' also possessed a notochord, along with seven pairs of pharyngeal bars, possibly made of cartilage. Surprisingly they were not formed from a singular bone, but they were formed of multiple separate pairs of bones, along with first two of them that were enlarged compared to t ...
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Metaspriggina
''Metaspriggina'' is a genus of chordate initially known from two specimens in the Middle Cambrian Burgess Shale and 44 specimens found in 2012 at the Marble Canyon bed in Kootenay National Park. Whilst named after the Ediacaran organism ''Spriggina'', later work has shown the two to be unrelated. ''Metaspriggina'' is considered to represent a primitive chordate, possibly transitional between cephalochordates and the earliest vertebrates, albeit this has been questioned because it seems to possess most of the characteristics attributed to craniates. It lacked fins and it had a weakly developed cranium, but it did possess two well-developed upward-facing eyes with nostrils behind them. ''Metaspriggina'' also possessed a notochord, along with seven pairs of pharyngeal bars, possibly made of cartilage. Surprisingly they were not formed from a singular bone, but they were formed of multiple separate pairs of bones, along with first two of them that were enlarged compared to t ...
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Metaspriggina NT Small
''Metaspriggina'' is a genus of chordate initially known from two specimens in the Middle Cambrian Burgess Shale and 44 specimens found in 2012 at the Marble Canyon bed in Kootenay National Park. Whilst named after the Ediacaran organism ''Spriggina'', later work has shown the two to be unrelated. ''Metaspriggina'' is considered to represent a primitive chordate, possibly transitional between cephalochordates and the earliest vertebrates, albeit this has been questioned because it seems to possess most of the characteristics attributed to craniates. It lacked fins and it had a weakly developed cranium, but it did possess two well-developed upward-facing eyes with nostrils behind them. ''Metaspriggina'' also possessed a notochord, along with seven pairs of pharyngeal bars, possibly made of cartilage. Surprisingly they were not formed from a singular bone, but they were formed of multiple separate pairs of bones, along with first two of them that were enlarged compared t ...
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Cambrian Chordate
The Cambrian chordates are an extinct group of animals belonging to the phylum Chordata that lived during the Cambrian, between 485 and 538 million years ago. The first Cambrian chordate known is '' Pikaia gracilens'', a lancelet-like animal from the Burgess Shale in British Columbia, Canada. The discoverer, Charles Doolittle Walcott, described it as a kind of worm (annelid) in 1911, but was later realised to be a chordate. Since the discovery of other Cambrian fossils from the Burgess Shale in 1991, and from the Chengjiang biota of China in 1991, which were later found to be of chordates, several Cambrian chordates are known, with some fossils considered as putative chordates. The Cambrian chordates are characterised by the presence of segmented muscle blocks called myomeres and notochord, the two defining features of chordates. Before the full understanding of Cambrian fossils, chordates as members the most advanced phylum were believed to appear on Earth much later than the Cam ...
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Marble Canyon (Canadian Rockies)
Marble Canyon is a canyon surrounding Tokumm Creek just above its confluence with the Vermilion River, at the north end of Kootenay National Park in the Canadian Rockies of British Columbia, Canada. South of Marble Canyon on Highway 93 is Numa Falls on the Vermillion River. As described by ''Canadian Alpine Journal'' in 1913, " okumm Creekjoins Vermillion River through a magnificent gorge, or box canyon, so narrow that at several places the fissure, for it seems little more than a crack in the rock strata, is bridged by great boulders that have become wedged across it. It was a feature well worth seeing." Cambrian Lagerstatte A major new find was announced in early 2014 of fossilized Cambrian soft-bodied organisms in or near Marble Canyon that rival or even surpass the nearby Burgess Shale fossil site in size and preservation. The report said that 22% of the observed species found in the initial excavation were new to science. Additionally, several species previously known o ...
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Pikaia
''Pikaia gracilens'' is an extinct, primitive chordate animal known from the Middle Cambrian Burgess Shale of British Columbia. Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became the "most famous early chordate fossil," or "famously known as the earliest" chordates. It is estimated to have lived during the latter period of the Cambrian explosion. Since it initial discovery, more than a hundred specimens have been recovered. The body structure resembles that of the lancelet and perhaps it swam much like an eel. A notochord and myomeres (segmented blocks of skeletal muscles) span the entire length of the body, and are considered as the defining signatures of chordate characters. Its primitive feature is indicated by the body covering, a cuticle, which is characteristic of invertebrates and some protochordates. The exact phylogenetic position is unclear. Proposed affinities include ...
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Chordate
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also Bilateral symmetry, bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit Metameric, metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cep ...
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Chordate
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also Bilateral symmetry, bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit Metameric, metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cep ...
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Nostril
A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbinates, whose function is to warm air on inhalation and remove moisture on exhalation. Fish do not breathe through noses, but they do have two (but cyclostomes have merged into one) small holes used for smelling, which can also be referred to as nostrils. In humans, the nasal cycle is the normal ultradian cycle of each nostril's blood vessels becoming engorged in swelling, then shrinking. The nostrils are separated by the septum. The septum can sometimes be deviated, causing one nostril to appear larger than the other. With extreme damage to the septum and columella, the two nostrils are no longer separated and form a single larger external opening. Like other tetrapods, humans have two external nostrils (anterior nares) and two additi ...
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Notochord
In anatomy, the notochord is a flexible rod which is similar in structure to the stiffer cartilage. If a species has a notochord at any stage of its life cycle (along with 4 other features), it is, by definition, a chordate. The notochord consists of inner, vacuolated cells covered by fibrous and elastic sheaths, lies along the anteroposterior axis (''front to back''), is usually closer to the dorsal than the ventral surface of the embryo, and is composed of cells derived from the mesoderm. The most commonly cited functions of the notochord are: as a midline tissue that provides directional signals to surrounding tissue during development, as a skeletal (structural) element, and as a vertebral precursor. In lancelets the notochord persists throughout life as the main structural support of the body. In tunicates the notochord is present only in the larval stage, being completely absent in the adult animal. In these invertebrate chordates, the notochord is not vacuolated. In all ...
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Cartilage
Cartilage is a resilient and smooth type of connective tissue. In tetrapods, it covers and protects the ends of long bones at the joints as articular cartilage, and is a structural component of many body parts including the rib cage, the neck and the bronchial tubes, and the intervertebral discs. In other taxa, such as chondrichthyans, but also in cyclostomes, it may constitute a much greater proportion of the skeleton. It is not as hard and rigid as bone, but it is much stiffer and much less flexible than muscle. The matrix of cartilage is made up of glycosaminoglycans, proteoglycans, collagen fibers and, sometimes, elastin. Because of its rigidity, cartilage often serves the purpose of holding tubes open in the body. Examples include the rings of the trachea, such as the cricoid cartilage and carina. Cartilage is composed of specialized cells called chondrocytes that produce a large amount of collagenous extracellular matrix, abundant ground substance that is rich in pro ...
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Seabed
The seabed (also known as the seafloor, sea floor, ocean floor, and ocean bottom) is the bottom of the ocean. All floors of the ocean are known as 'seabeds'. The structure of the seabed of the global ocean is governed by plate tectonics. Most of the ocean is very deep, where the seabed is known as the abyssal plain. Seafloor spreading creates mid-ocean ridges along the center line of major ocean basins, where the seabed is slightly shallower than the surrounding abyssal plain. From the abyssal plain, the seabed slopes upward toward the continents and becomes, in order from deep to shallow, the continental rise, slope, and shelf. The depth within the seabed itself, such as the depth down through a sediment core, is known as the “depth below seafloor.” The ecological environment of the seabed and the deepest waters are collectively known, as a habitat for creatures, as the “benthos.” Most of the seabed throughout the world's oceans is covered in layers of marine sediments. ...
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Gnathostomata
Gnathostomata (; from Greek: (') "jaw" + (') "mouth") are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. In addition to opposing jaws, living gnathostomes have true teeth (a characteristic which has subsequently been lost in some), paired appendages (pectoral and pelvic fins, arms, legs, wings, etc.), the elastomeric protein of elastin, and a horizontal semicircular canal of the inner ear, along with physiological and cellular anatomical characters such as the myelin sheaths of neurons, and an adaptive immune system that has the discrete lymphoid organs of spleen and thymus, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene. It is now assumed that Gnathostomata evolved from ancestors that already possessed a pair of both pectoral and pelvic fins. Until recently these ancestors, know ...
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