Megadolodus
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Megadolodus
''Megadolodus'' is an extinct genus of proterotheriid litopterns. History The first fossils of ''Megadolodus'' were uncovered from the Villavieja Formation, in the fossil locality La Venta, in what is today Colombia, in terrains dated from the Middle Miocene, between 13.5 and 11.8 millions of years ago. The genus was described in 1956 after the holotype UCMP 39270, consisting in a left mandible preserving the fourth premolar, the first molar and roots from the second molar, and discovered during an expedition organized by the University of California and directed by Ruben A. Stirton. Those few remains led Malcolm McKenna,Malcolm C. McKenna. Survival of primitive notoungulates and condylarths into the Miocene of Colombia. ''American Journal of Science'', Vol. 254, December 1956, P.736-743 who described them in his 1956 article, to believe they originated from a late surviving Didolodontidae, a family of primitive ungulates that went extinct during the end of the Eocene, ...
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Laventan
The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes the Mayoan age.Madden et al., 1997 Etymology The age is named after the Miocene Lagerstätte La Venta, where a rich biodiversity from the Middle Miocene has been recovered from the Honda Group. Formations Fossil content Correlations The Laventan (13.8 to 11.8 Ma) correlates with: * NALMA ** latest Barstovian (15.97-13.65 Ma)Barstovian
at Fossilworks.org
** early

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Proterotheriidae
Proterotheriidae is an extinction, extinct family of fossil ungulates from the Cenozoic era that displays toe reduction. Despite resembling primitive, small horses, they were only distantly related to them, and instead belonged to the native South American ungulate order Litopterna. Description Two subfamilies and 18 genera of Proterotheriidae are known. All forms were small or medium-sized. Typical is a reduction of the number of toes and brachydont or mesodont teeth. The family is recorded since the late Palaeocene. Various fossils are known from many parts of the South American continent. The diversity decreased in the Miocene to Pliocene and it has been assumed for a long time that they entirely disappeared in the late Pliocene. However, fossils found in Argentina, Brazil and Uruguay show that one member of the group, ''Neolicaphrium recens'' survived into the Late Lujanian, Pleistocene. Better known genera of the family include ''Diadiaphorus'' and ''Thoatherium'' from th ...
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Honda Group, Colombia
The Honda Group ( es, Grupo Honda, Tsh, Ngh) is a geological group of the Upper and Middle Magdalena Basins and the adjacent Central and Eastern Ranges of the Colombian Andes. The group, in older literature also defined as formation, is in its present-day type section in the Tatacoa Desert in the department of Huila subdivided into two main formations; La Victoria and Villavieja. The group was originally defined in and named after Honda, Tolima, but has been redefined based on the many fossil finds in the Tatacoa Desert, to the south. In the original type section of its occurrence, the thick group is subdivided into three formations, from old to young; Cambrás, San Antonio and Los Limones. The group dates to the Neogene period; in its broadest definition from the Late Oligocene to Late Miocene, and in the redefined type section restricted to the Laventan age of the South American Land Mammal Ages (SALMA), equivalent to the Middle Miocene Serravallian epoch. The Honda Gr ...
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La Venta (Colombia)
La Venta is a fossil locality located in the modern departments of Tolima and Huila in Colombia. This site is one of the richest Neogene fossil assemblages in South America and represents the best-known Cenozoic fossil site outside of Argentina. It provides a glimpse of what life in the region was like before the main wave of the Great American Interchange. Geology The fossils occur in Middle Miocene rocks of the Honda Group, which is divided into the younger Villavieja Formation and the older La Victoria Formation. The La Venta fauna contained ancient species of animal genera and families still alive today, as well as some entirely extinct prehistoric lineages. These animals lived some 13.8 to 12 million years ago in the Laventan age, which was named after La Venta. At that time, the climate of the region was wetter than today and there was much forest of trees similar (and probably related) to the ''sapino'' (''Goupia glabra'') of our time. List of fauna Fossil animals f ...
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Bounodus
''Bounodus'' is an extinct genus of ungulates native from South America, belonging to the order Litopterna. Its remains were found in Miocene rocks from the Urumaco Formation of northern Venezuela. The genus is only known from its holotype, AMU-CURS 44, a fragmentary right maxilla preserving teeth from the fourth premolar to the third molar and the alveolus of the third premolar, exhibiting a typical bunodont structure in the animal dentition, a characteristic shared with its close relative ''Megadolodus''. The postcranial skeleton of the genus is unknown. Its dentition indicates that the animal was omnivorous, with teeth adapted to potentially eat shelled fruits, as well as the molluscs and crustaceans that thrived in the wetlands that covered most of North-West South America, including the Urumaco Formation, during the Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of ...
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Diadiaphorus
''Diadiaphorus'' is an extinct genus of litoptern mammal from the Miocene of Argentina ( Ituzaingó, Pinturas, Chiquimil and Santa Cruz Formations) and Bolivia ( Nazareno Formation), South America. Description ''Diadiaphorus'' closely resembled a horse, but was only around in body length with a weight , similar to a modern sheep.D. Patterson, Bruce (2012) ''Bones, Clones, and Biomes: The History and Geography of Recent Neotropical Mammals'' p.92 It had three toes, only one of which touched the ground. This toe had a large hoof; the two outer toes were rudimentary, much like those of early horses such as ''Merychippus''. Unlike horses, however, ''Diadiaphorus'' lacked fused limb bones. Its skull was short and had a relatively large brain cavity. Judging from its low molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from La ...
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Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in the Caribbean Sea. It has a territorial extension of , and its population was estimated at 29 million in 2022. The capital and largest urban agglomeration is the city of Caracas. The continental territory is bordered on the north by the Caribbean Sea and the Atlantic Ocean, on the west by Colombia, Brazil on the south, Trinidad and Tobago to the north-east and on the east by Guyana. The Venezuelan government maintains a claim against Guyana to Guayana Esequiba. Venezuela is a federal presidential republic consisting of 23 states, the Capital District and federal dependencies covering Venezuela's offshore islands. Venezuela is among the most urbanized countries in Latin America; the vast majority of Venezuelans live in the cities of the n ...
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Horses
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature, ''Eohippus'', into the large, single-toed animal of today. Humans began domesticating horses around 4000 BCE, and their domestication is believed to have been widespread by 3000 BCE. Horses in the subspecies ''caballus'' are domesticated, although some domesticated populations live in the wild as feral horses. These feral populations are not true wild horses, as this term is used to describe horses that have never been domesticated. There is an extensive, specialized vocabulary used to describe equine-related concepts, covering everything from anatomy to life stages, size, colors, markings, breeds, locomotion, and behavior. Horses are adapted to run, allowing them to quickly escape predators, and poss ...
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Thoatherium
''Thoatherium'' (meaning "active swift-beast") is an extinct genus of litoptern mammals from the Early Miocene of Argentina. Fossils of the genus have been found in the Santa Cruz Formation in Argentina.''Thoatherium''
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Description

With a length of , the -like ''Thoatherium'' was a small representative of the order
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Tusk
Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share common features such as extra-oral position, growth pattern, composition and structure, and lack of contribution to ingestion. Tusks are thought to have adapted to the extra-oral environments, like dry or aquatic or arctic. In most tusked species both the males and the females have tusks although the males' are larger. Most mammals with tusks have a pair of them growing out from either side of the mouth. Tusks are generally curved and have a smooth, continuous surface. The male narwhal's straight single helical tusk, which usually grows out from the left of the mouth, is an exception to the typical features of tusks described above. Continuous growth of tusks is enabled by formative tissues in the apical openings of the roots of the teeth. ...
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