Marada (mammal)
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Marada (mammal)
''Marada arcanum'' is a species of Vombatiformes discovered in 2001 at the Oligocene Hiatus Site at Riversleigh. It is the only member of the genus ''Marada''. It exhibits plesiomorphic and apomorphic features making the determination of its taxonomic placement difficult. It has been placed within its own family, Maradidae. The specimen consists of the right dentary, with the first incisor but missing the crown, the whole of the horizontal ramus with intact premolar three and molars one to four. The posterior is missing the coronoid process, the articular condyle and the angular process. The new species, genus, and family were described in 2007 by the Australian palaeontologist Karen Black Karen Blanche Black (née Ziegler; July 1, 1939 – August 8, 2013) was an American actress, screenwriter, singer, and songwriter. She rose to prominence for her work in various studio and independent films in the 1970s, frequently portrayi ... References {{Taxonbar, from1=Q6540 ...
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Upper Oligocene
The Chattian is, in the geologic timescale, the younger of two ages or upper of two stages of the Oligocene Epoch/Series. It spans the time between . The Chattian is preceded by the Rupelian and is followed by the Aquitanian (the lowest stage of the Miocene). Stratigraphic definition The Chattian was introduced by Austrian palaeontologist Theodor Fuchs in 1894. Fuchs named the stage after the Chatti, a Germanic tribe.Berry, Edward W"The Mayence Basin, a Chapter of Geologic History" ''The Scientific Monthly'', Vol. 16, No. 2, February 1923. pp. 114. Retrieved March 18, 2020. The original type locality was near the German city of Kassel. The base of the Chattian is at the extinction of the foram genus ''Chiloguembelina'' (which is also the base of foram biozone P21b). An official GSSP for the Chattian Stage was ratified in October of 2016. The top of the Chattian Stage (which is the base of the Aquitanian Stage, Miocene Series and Neogene System) is at the first appearance of ...
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Karen Black (palaeontologist)
Karen H. Black, born about 1970, is a palaeontologist at the University of New South Wales. Black is the leading author on research describing new families, genera and species of fossil mammals. She is interested in understanding faunal change and community structure in order to gain new understandings of past, current and future changes in biodiversity which are driven by climate. Karen Black won the Dorothy Hill medal, from the Australian Academy of Science in 2012, for research on the genus ''Nimbadon'', and is recognised by fellow researchers in the specific epithet of ''Hypsiprymnodon karenblackae''. Career Black's early career involved extracting, curating and analysing the fossils of the vertebrate faunas within limestone deposits in the region of the Riversleigh World Heritage Area, located in northwestern Queensland. Black's research is focussed on improving our understanding of species interactions, development, faunal change, as well as behaviour, and community s ...
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Vombatiformes
The Vombatiformes are one of the three suborders of the large marsupial order Diprotodontia. Seven of the nine known families within this suborder are extinct; only the families Phascolarctidae, with the koala, and Vombatidae, with three extant species of wombat, survive. Among the extinct families are the Diprotodontidae, which includes the rhinoceros sized ''Diprotodon'', believed to be the largest marsupials ever, as well as the "marsupial lions" Thylacoleonidae and "marsupial tapirs" Palorchestidae. "Vombatiformes" is neo-Latin for "wombat-shaped things", and took its name from its type family. The suborder Vombatiformes, with its closely related members and their compact body form, contrasts with the other two diprotodont suborders, the Macropodiformes: kangaroos, wallabies, and the quokka; and the Phalangeriformes: possums, including the gliders such as the wrist-winged gliders. The koala and wombats are believed by many biologists to share a common ancestor and to have di ...
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Australian Fossil Mammal Sites (Riversleigh)
Riversleigh World Heritage Area is Australia's most famous fossil location, recognised for the series of well preserved fossils deposited from the Late Oligocene to more recent geological periods. The fossiliferous limestone system is located near the Gregory River in the north-west of Queensland, an environment that was once a very wet rainforest that became more arid as the Gondwanan land masses separated and the Australian continent moved north. The approximately area has fossil remains of ancient mammals, birds, and reptiles of the Oligocene and Miocene ages, many of which were discovered and are only known from the Riversleigh area; the species that have occurred there are known as the Riversleigh fauna. The fossils at Riversleigh are unusual because they are found in soft freshwater limestone which has not been compacted. This means the animal remains retain their three-dimensional structure, rather than being partially crushed like in most fossil sites. The area is loca ...
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Plesiomorphic
In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and synapomorphy, all mean a trait shared between species because they share an ancestral species. Apomorphic and synapomorphic characteristics convey much information about evolutionary clades and can be used to define taxa. However, plesiomorphic and symplesiomorphic characteristics cannot. The term ''symplesiomorphy'' was introduced in 1950 by German entomologist Willi Hennig. Examples A backbone is a plesiomorphic trait shared by birds and mammals, and does not help in placing an animal in one or the other of these two clades. Birds and mammals share this trait because both clades are descended from the same far distant ancestor. Other clades, e.g. snakes, lizards, turtles, fish, frogs, all have backbones and none are either birds no ...
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Apomorphic
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis Th ...
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Incisor
Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whereas armadillos have none. Structure Adult humans normally have eight incisors, two of each type. The types of incisor are: * maxillary central incisor (upper jaw, closest to the center of the lips) * maxillary lateral incisor (upper jaw, beside the maxillary central incisor) * mandibular central incisor (lower jaw, closest to the center of the lips) * mandibular lateral incisor (lower jaw, beside the mandibular central incisor) Children with a full set of deciduous teeth (primary teeth) also have eight incisors, named the same way as in permanent teeth. Young children may have from zero to eight incisors depending on the stage of their tooth eruption and tooth development. Typically, the mandibular central incisors erupt first, followed ...
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Crown (tooth)
In dentistry, crown refers to the anatomical area of teeth, usually covered by enamel. The crown is usually visible in the mouth after developing below the gingiva The gums or gingiva (plural: ''gingivae'') consist of the mucosal tissue that lies over the mandible and maxilla inside the mouth. Gum health and disease can have an effect on general health. Structure The gums are part of the soft tissue lin ... and then erupting into place. If part of the tooth gets chipped or broken, a dentist can apply an artificial crown. Crowns are used most commonly to entirely cover a damaged tooth or cover an implant. Bridges are also used to cover a space if one or more teeth is missing. They are cemented to natural teeth or implants surrounding the space where the tooth once stood. There are various materials that can be used including a type of cement or stainless steel. The cement crowns look like regular teeth while the stainless steel crowns are silver or gold. References ...
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Premolar
The premolars, also called premolar teeth, or bicuspids, are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant in the permanent set of teeth, making eight premolars total in the mouth. They have at least two cusps. Premolars can be considered transitional teeth during chewing, or mastication. They have properties of both the canines, that lie anterior and molars that lie posterior, and so food can be transferred from the canines to the premolars and finally to the molars for grinding, instead of directly from the canines to the molars. Human anatomy The premolars in humans are the maxillary first premolar, maxillary second premolar, mandibular first premolar, and the mandibular second premolar. Premolar teeth by definition are permanent teeth distal to the canines, preceded by deciduous molars. Morphology There is always one large buccal cusp, especially so in the mandibular first premolar. The lower second ...
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Molar (tooth)
The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone tooth", from ''mola'', millstone and ''dens'', tooth. Molars show a great deal of diversity in size and shape across mammal groups. The third molar of humans is sometimes vestigial. Human anatomy In humans, the molar teeth have either four or five cusps. Adult humans have 12 molars, in four groups of three at the back of the mouth. The third, rearmost molar in each group is called a wisdom tooth. It is the last tooth to appear, breaking through the front of the gum at about the age of 20, although this varies from individual to individual. Race can also affect the age at which this occurs, with statistical variations between groups. In some cases, it may not even erupt at all. The human mouth contains upper (maxillary) and lower (mandib ...
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Coronoid Process Of The Mandible
In human anatomy, the mandible's coronoid process (from Greek ''korōnē'', denoting something hooked) is a thin, triangular eminence, which is flattened from side to side and varies in shape and size. Its anterior border is convex and is continuous below with the anterior border of the ramus. Its ''posterior border'' is concave and forms the anterior boundary of the mandibular notch. The ''lateral surface'' is smooth, and affords insertion to the temporalis and masseter muscles. Its ''medial surface'' gives insertion to the temporalis, and presents a ridge which begins near the apex of the process and runs downward and forward to the inner side of the last molar tooth. Between this ridge and the anterior border is a grooved triangular area, the upper part of which gives attachment to the temporalis, the lower part to some fibers of the buccinator. Clinical significance Fractures of the mandible are common. However, coronoid process fractures are very rare. Isolated fractures of th ...
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Prehistoric Vombatiforms
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. ...
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