Macivera
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Macivera
''Macivera'' is a genus of extinct vascular plants. Fossils were found in sediments in Bathust Island, Nunavut, Canada, from the upper Silurian (Ludfordian, around ). The leafless stems (axes) branched dichotomously and were relatively thin, being between 0.7 and 1.0 mm wide. Spore-forming organs or sporangia, which were elliptical, being longer than wide, were borne on the end regions of stems. ''Macivera'' is considered to be a zosterophyll. Description The genus was first described from a small number of specimens found in sediments in Bathurst Island, Nunavut, Canada, which are considered to be of Late Silurian age (Ludfordian, around ). Stems (axes) were smooth, devoid of leaves, hairs or other protrusions, and were between 0.7 and 1.0 mm wide (around the lower limit of the size Boyce suggested as being compatible with the presence of vascular tissue), tapering down to 0.1 mm towards the apex of those stems which did not bear sporangia. The stems branched eq ...
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Macivera
''Macivera'' is a genus of extinct vascular plants. Fossils were found in sediments in Bathust Island, Nunavut, Canada, from the upper Silurian (Ludfordian, around ). The leafless stems (axes) branched dichotomously and were relatively thin, being between 0.7 and 1.0 mm wide. Spore-forming organs or sporangia, which were elliptical, being longer than wide, were borne on the end regions of stems. ''Macivera'' is considered to be a zosterophyll. Description The genus was first described from a small number of specimens found in sediments in Bathurst Island, Nunavut, Canada, which are considered to be of Late Silurian age (Ludfordian, around ). Stems (axes) were smooth, devoid of leaves, hairs or other protrusions, and were between 0.7 and 1.0 mm wide (around the lower limit of the size Boyce suggested as being compatible with the presence of vascular tissue), tapering down to 0.1 mm towards the apex of those stems which did not bear sporangia. The stems branched eq ...
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Zosterophyll
The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosterophyllophyta or Zosterophyta and the class or plesion Zosterophyllopsida or Zosteropsida. They were among the first vascular plants in the fossil record, and had a world-wide distribution. They were probably stem-group lycophytes, forming a sister group to the ancestors of the living lycophytes. By the late Silurian (late Ludlovian, about ) a diverse assemblage of species existed, examples of which have been found fossilised in what is now Bathurst Island in Arctic Canada. Morphology The stems of zosterophylls were either smooth or covered with small spines known as enations, branched dichotomously, and grew at the ends by unrolling, a process known as circinate vernation. The stems had a central vascular column in which the protoxylem ...
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Zosterophylls
The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosterophyllophyta or Zosterophyta and the class or plesion Zosterophyllopsida or Zosteropsida. They were among the first vascular plants in the fossil record, and had a world-wide distribution. They were probably stem-group lycophytes, forming a sister group to the ancestors of the living lycophytes. By the late Silurian (late Ludlovian, about ) a diverse assemblage of species existed, examples of which have been found fossilised in what is now Bathurst Island in Arctic Canada. Morphology The stems of zosterophylls were either smooth or covered with small spines known as enations, branched dichotomously, and grew at the ends by unrolling, a process known as circinate vernation. The stems had a central vascular column in which the protoxylem ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Vascular Plant
Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They also have a specialized non-lignified tissue (the phloem) to conduct products of photosynthesis. Vascular plants include the clubmosses, horsetails, ferns, gymnosperms (including conifers), and angiosperms (flowering plants). Scientific names for the group include Tracheophyta, Tracheobionta and Equisetopsida ''sensu lato''. Some early land plants (the rhyniophytes) had less developed vascular tissue; the term eutracheophyte has been used for all other vascular plants, including all living ones. Historically, vascular plants were known as "higher plants", as it was believed that they were further evolved than other plants due to being more complex organisms. However, this is an antiquated remnant of the obsolete scala naturae, and the term ...
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Ludfordian
In the geologic timescale, the Ludfordian is the upper of two chronostratigraphic stages within the Ludlow Series. Its age is the late Silurian Period, and within both the Palaeozoic Era and Phanerozoic Eon. The rocks assigned to the Ludfordian date to between 425.6 ± 0.9 Ma and 423.0 ± 2.3 Ma (million years ago). The Ludfordian Stage succeeds the Gorstian Stage and precedes the Pridoli Epoch. It is named for the village of Ludford in Shropshire, England. The GSSP for the Ludfordian is represented as a thin shale seam, coincident with the base of the Leintwardine Formation, overlying the Bringewood Formation in England. Paleoclimate The Lau event is a rapid pulse of cooling during the Ludfordian, about ; it is identified by a pulse of extinctions and oceanic changes. It is one of the series of fast sea-level and excursions in oxygen isotope ratios that signal fast switches between warm and cold climate states, characteristic of the Silurian climatic instability. The Lau ...
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Spore
In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. Under favourable conditions the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote which develops into a new s ...
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Sporangium
A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cycle. Sporangia can produce spores by mitosis, but in nearly all land plants and many fungi, sporangia are the site of meiosis and produce genetically distinct haploid spores. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains haploid nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and germinate into haploid hyphae. Although sexual reproduction in fungi varies between phyla, for some fungi the sporangium plays an indirect role in sexual reprod ...
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Late Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during t ...
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Vascular Tissue
Vascular tissue is a complex conducting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and phloem. These two tissues transport fluid and nutrients internally. There are also two meristems associated with vascular tissue: the vascular cambium and the cork cambium. All the vascular tissues within a particular plant together constitute the vascular tissue system of that plant. The cells in vascular tissue are typically long and slender. Since the xylem and phloem function in the conduction of water, minerals, and nutrients throughout the plant, it is not surprising that their form should be similar to pipes. The individual cells of phloem are connected end-to-end, just as the sections of a pipe might be. As the plant grows, new vascular tissue differentiates in the growing tips of the plant. The new tissue is aligned with existing vascular tissue, maintaining its connection throughout the plant. The ...
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Paleobotanist
Paleobotany, which is also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeography), and the evolutionary history of plants, with a bearing upon the evolution of life in general. A synonym is paleophytology. It is a component of paleontology and paleobiology. The prefix ''palaeo-'' means "ancient, old", and is derived from the Greek adjective , . Paleobotany includes the study of terrestrial plant fossils, as well as the study of prehistoric marine photoautotrophs, such as photosynthetic algae, seaweeds or kelp. A closely related field is palynology, which is the study of fossilized and extant spores and pollen. Paleobotany is important in the reconstruction of ancient ecological systems and climate, known as paleoecology and paleoclimatology respectively; and is fundamental to the study of green plant developm ...
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Distichophytum
''Distichophytum'' is a genus of extinct vascular plants of the Late Silurian (Ludfordian) to Early Devonian (Emsian), around . The genus has a tangled taxonomic history, also being known as ''Bucheria'' and ''Rebuchia'' (see below). Description The genus was first discovered as fossils of Early Devonian age (Pragian or Siegenian to Emsian, ), consisting of isolated spikes of sporangia (spore-forming organs) found at Beartooth Butte, Wyoming, United States of America. Specimens of ''D. ovata'' with sporangia attached to stems were later found at the same location. The base of the plant remains unknown; the known part was about 8.5 cm high. The sporophyte consisted of narrow leafless stems (axes) 1.5 to 2.0 mm in diameter, which branched dichotomously. Stems which did not bear sporangia ended in blunt points; fertile branches bore compact one-sided spikes of up to 20 laterally attached sporangia, more-or-less opposite. The sporangia were kidney-shaped (reniform) and ha ...
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