Lonchognatha
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Lonchognatha
Novialoidea (meaning "new wings") is an extinct clade of macronychopteran pterosaurs that lived from the latest Early Jurassic to the latest Late Cretaceous (early Toarcian to late Maastrichtian ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107/ref>), their fossils having been found on all continents except Antarctica. History and classification Novialoidea was named by paleontologist Alexander Wilhelm Armin Kellner in 2003 as a node-based taxon consisting of the last common ancestor of ''Campylognathoides'', '' Quetzalcoatlus'' and all its descendants. This name was derived from Latin ''novus'' "new", and ''ala'', "wing", in reference to the wing synapomorphies that the members of the clade possess.Kellner, A. W. A., (2003): Pterosaur phylogeny and comments on the evolutionary history of the group. pp. 105-137. — ''in'' Buffetaut, E. & Mazin, J.-M., (eds.): ''Evolution an ...
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Campylognathoides Liasicus
''Campylognathoides'' ("curved jaw", Strand 1928) is an extinct genus of pterosaur discovered in the Württemberg Lias deposits (dated to the early Toarcian ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). "Pterosaur distribution in time and space: an atlas". ''Zitteliana'', 61–107/ref>) of Germany; this first specimen however, consisted only of wing fragments. Further better preserved specimens were found in the Holzmaden shale; based on these specimens, Felix Plieninger erected a new genus. Discovery In 1858 Friedrich August Quenstedt named a new species of ''Pterodactylus'': ''P. liasicus''. It was based on a fossil, holotype GPIT 9533, consisting of some wing bones, found on the Wittberg near Metzingen in layers dating from the early Toarcian, about 180 million years old. The specific name referred to the Lias. Quenstedt thought he had identified long metacarpals in the wing, concluding that the new species was therefore not belonging to more basa ...
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Eudimorphodon
''Eudimorphodon'' was a pterosaur that was discovered in 1973 by Mario Pandolfi in the town of Cene, Italy and described the same year by Rocco Zambelli. The nearly complete skeleton was retrieved from shale deposited during the Late Triassic (mid to late Norian stage),Wellnhofer, P. (1991). "Summary of Triassic Pterosaurs." ''The Illustrated Encyclopedia of Pterosaurs.'' London, UK: Salamander Books Limited. p. 67. . making ''Eudimorphodon'' one of the oldest pterosaurs known.Cranfield, I. ''The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures''. London: Salamander Books, Ltd. Pp. 280–281. It had a wingspan of about and at the end of its long bony tail may have been a diamond-shaped flap like in the later ''Rhamphorhynchus''. If so, the flap may have helped it steer while maneuvering in the air. ''Eudimorphodon'' is known from several skeletons, including juvenile specimens. Discovery and species ''Eudimorphodon'' currently includes one species, the type sp ...
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Campylognathoides
''Campylognathoides'' ("curved jaw", Strand 1928) is an extinct genus of pterosaur discovered in the Württemberg Lias deposits (dated to the early Toarcian ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). "Pterosaur distribution in time and space: an atlas". ''Zitteliana'', 61–107/ref>) of Germany; this first specimen however, consisted only of wing fragments. Further better preserved specimens were found in the Holzmaden shale; based on these specimens, Felix Plieninger erected a new genus. Discovery In 1858 Friedrich August Quenstedt named a new species of ''Pterodactylus'': ''P. liasicus''. It was based on a fossil, holotype GPIT 9533, consisting of some wing bones, found on the Wittberg near Metzingen in layers dating from the early Toarcian, about 180 million years old. The specific name referred to the Lias. Quenstedt thought he had identified long metacarpals in the wing, concluding that the new species was therefore not belonging to more bas ...
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Eudimorphodon Ranzii
''Eudimorphodon'' was a pterosaur that was discovered in 1973 by Mario Pandolfi in the town of Cene, Italy and described the same year by Rocco Zambelli. The nearly complete skeleton was retrieved from shale deposited during the Late Triassic (mid to late Norian stage),Wellnhofer, P. (1991). "Summary of Triassic Pterosaurs." ''The Illustrated Encyclopedia of Pterosaurs.'' London, UK: Salamander Books Limited. p. 67. . making ''Eudimorphodon'' one of the oldest pterosaurs known.Cranfield, I. ''The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures''. London: Salamander Books, Ltd. Pp. 280–281. It had a wingspan of about and at the end of its long bony tail may have been a diamond-shaped flap like in the later ''Rhamphorhynchus''. If so, the flap may have helped it steer while maneuvering in the air. ''Eudimorphodon'' is known from several skeletons, including juvenile specimens. Discovery and species ''Eudimorphodon'' currently includes one species, the type sp ...
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Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', t ...
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Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary extinction event) occurred at the end of this age. In this mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous. Stratigraphic definitions Definition The Maastrichtian was introduced into scientific literature by Belgian geologist André Hubert Dumont in 1849, after studying rock strata of the Chalk Group c ...
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Rhamphorhynchus
''Rhamphorhynchus'' (, from Ancient Greek ''rhamphos'' meaning "beak" and ''rhynchus'' meaning "snout") is a genus of long-tailed pterosaurs in the Jurassic period. Less specialized than contemporary, short-tailed pterodactyloid pterosaurs such as ''Pterodactylus'', it had a long tail, stiffened with ligaments, which ended in a characteristic soft-tissue tail vane. The mouth of ''Rhamphorhynchus'' housed needle-like teeth, which were angled forward, with a curved, sharp, beak-like tip lacking teeth, indicating a diet mainly of fish; indeed, fish and cephalopod remains are frequently found in ''Rhamphorhynchus'' abdominal contents, as well as in their coprolites. Although fragmentary fossil remains possibly belonging to ''Rhamphorhynchus'' have been found in England, Tanzania, and Spain, the best preserved specimens come from the Solnhofen limestone of Bavaria, Germany. Many of these fossils preserve not only the bones but impressions of soft tissues, such as wing membranes. Scat ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Rhamphorhynchus Muensteri
''Rhamphorhynchus'' (, from Ancient Greek ''rhamphos'' meaning "beak" and ''rhynchus'' meaning "snout") is a genus of long-tailed pterosaurs in the Jurassic period. Less specialized than contemporary, short-tailed pterodactyloid pterosaurs such as '' Pterodactylus'', it had a long tail, stiffened with ligaments, which ended in a characteristic soft-tissue tail vane. The mouth of ''Rhamphorhynchus'' housed needle-like teeth, which were angled forward, with a curved, sharp, beak-like tip lacking teeth, indicating a diet mainly of fish; indeed, fish and cephalopod A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head ... remains are frequently found in ''Rhamphorhynchus'' abdominal contents, as well as in their coprolites. Although fragmentary fossil remains possibly belonging to ''Rhampho ...
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Geological Society Of London
The Geological Society of London, known commonly as the Geological Society, is a learned society based in the United Kingdom. It is the oldest national geological society in the world and the largest in Europe with more than 12,000 Fellows. Fellows are entitled to the postnominal FGS (Fellow of the Geological Society), over 2,000 of whom are Chartered Geologists (CGeol). The Society is a Registered Charity, No. 210161. It is also a member of the Science Council, and is licensed to award Chartered Scientist to qualifying members. The mission of the society is: "Making geologists acquainted with each other, stimulating their zeal, inducing them to adopt one nomenclature, facilitating the communication of new facts and ascertaining what is known in their science and what remains to be discovered". History The Society was founded on 13 November 1807 at the Freemasons' Tavern, Great Queen Street, in the Covent Garden district of London. It was partly the outcome of a previous cl ...
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David Unwin
David (; , "beloved one") (traditional spelling), , ''Dāwūd''; grc-koi, Δαυΐδ, Dauíd; la, Davidus, David; gez , ዳዊት, ''Dawit''; xcl, Դաւիթ, ''Dawitʿ''; cu, Давíдъ, ''Davidŭ''; possibly meaning "beloved one". was, according to the Hebrew Bible, the third king of the United Kingdom of Israel. In the Books of Samuel, he is described as a young shepherd and harpist who gains fame by slaying Goliath, a champion of the Philistines, in southern Canaan. David becomes a favourite of Saul, the first king of Israel; he also forges a notably close friendship with Jonathan, a son of Saul. However, under the paranoia that David is seeking to usurp the throne, Saul attempts to kill David, forcing the latter to go into hiding and effectively operate as a fugitive for several years. After Saul and Jonathan are both killed in battle against the Philistines, a 30-year-old David is anointed king over all of Israel and Judah. Following his rise to power, David c ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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