Letharia Columbiana
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Letharia Columbiana
''Letharia columbiana ''(common name brown-eye wolf lichen, synonyms ''Letharia californica'', ''Borrera columbiana'') is a common lichen in subalpine forests, particularly in the Pacific Northwest of the United States, and parts of Canada.  It is in the family Parmeliaceae, and the genus '' Letharia''.  Its characteristics include a bright citron color, “brown-eyes”, and rounded, irregular branches. Though previously believed to lump together several lineages such as ''Letharia gracilis'' and others, there now exists more specific characteristics to identify the species.  This lichen grows on the bark of conifers a couple inches tall.  ''L. Columbiana''’s cousin, ''Letharia vulpina ''(common name wolf lichen), has similar geographical distribution and morphological features, with the major difference being the “brown-eyes” of ''L. columbiana''. Habitat and distribution ''L. columbiana'' has a healthy distribution in subalpine forests of the United States and Canad ...
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Lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (



Parmeliaceae
The Parmeliaceae is a large and diverse family of Lecanoromycetes. With over 2700 species in 71 genera, it is the largest family of lichen-forming fungi. The most speciose genera in the family are the well-known groups: '' Xanthoparmelia'' ( 822 species), ''Usnea'' (355 species), ''Parmotrema'' ( 255 species), and ''Hypotrachyna'' (262 species). Nearly all members of the family have a symbiotic association with a green alga (most often ''Trebouxia'' spp., but '' Asterochloris'' spp. are known to associate with some species).Miadlikowska, J. ''et al.'' (2006). New insights into classification and evolution of the Lecanoromycetes (Pezizomycotina, Ascomycota) from phylogenetic analyses of three ribosomal RNA- and two protein-coding genes. ''Mycologia'' 98: 1088-1103. http://www.mycologia.org/cgi/reprint/98/6/1088.pdf The majority of Parmeliaceae species have a foliose, fruticose, or subfruticose growth form. The morphological diversity and complexity exhibited by this group is en ...
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Letharia
''Letharia'' is a genus of fruticose lichens belonging to the family Parmeliaceae. There were historically two species of ''Letharia'': '' L. vulpina'' and '' L. columbiana''. Recent molecular sequence studies published in 2016 confirm at least 6 species in Western North America alone, with more expected to be confirmed using similar methods in other parts of the world. The typical photobiont is a green alga of genus ''Trebouxia ''Trebouxia'' is a unicellular green alga.Silverside, A. J. (2009). Retrieved from http://www.bioref.lastdragon.org/Chlorophyta/''Trebouxia''.html It is a photosynthetic organism that can exist in almost all habitats found in polar, tropical, and ...''. References Parmeliaceae Lecanorales genera Lichen genera Taxa named by Theodor Magnus Fries Taxa described in 1871 {{Parmeliaceae-stub ...
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Conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class (biology), class, Pinopsida. All Neontology, extant conifers are perennial plant, perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include Cedrus, cedars, Pseudotsuga, Douglas-firs, Cupressaceae, cypresses, firs, junipers, Agathis, kauri, larches, pines, Tsuga, hemlocks, Sequoioideae, redwoods, spruces, and Taxaceae, yews.Campbell, Reece, "Phylum Coniferophyta". Biology. 7th. 2005. Print. P. 595 As of 1998, the division Pinophyta was estimated to contain eight families, 68 genera, and 629 living species. Although the total number of species is relatively small, conifers are ecology, ecologically important. They are the dominant plants over large areas of land, most ...
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Letharia Vulpina JHollinger Crop
''Letharia'' is a genus of fruticose lichens belonging to the family Parmeliaceae. There were historically two species of ''Letharia'': '' L. vulpina'' and '' L. columbiana''. Recent molecular sequence studies published in 2016 confirm at least 6 species in Western North America alone, with more expected to be confirmed using similar methods in other parts of the world. The typical photobiont is a green alga of genus ''Trebouxia ''Trebouxia'' is a unicellular green alga.Silverside, A. J. (2009). Retrieved from http://www.bioref.lastdragon.org/Chlorophyta/''Trebouxia''.html It is a photosynthetic organism that can exist in almost all habitats found in polar, tropical, a ...''. References Parmeliaceae Lecanorales genera Lichen genera Taxa named by Theodor Magnus Fries Taxa described in 1871 {{Parmeliaceae-stub ...
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Ascomycete
Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The defining feature of this fungal group is the " ascus" (), a microscopic sexual structure in which nonmotile spores, called ascospores, are formed. However, some species of the Ascomycota are asexual, meaning that they do not have a sexual cycle and thus do not form asci or ascospores. Familiar examples of sac fungi include morels, truffles, brewers' and bakers' yeast, dead man's fingers, and cup fungi. The fungal symbionts in the majority of lichens (loosely termed "ascolichens") such as ''Cladonia'' belong to the Ascomycota. Ascomycota is a monophyletic group (it contains all descendants of one common ancestor). Previously placed in the Deuteromycota along with asexual species from other fungal taxa, asexual (or anamorphic) ascomycetes ...
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Vulpicida Canadensis
''Vulpicida'' is a genus of lichenized fungi in the family Parmeliaceae. Circumscribed in 1993 to contain species formerly placed in ''Cetraria'', the genus is widespread in Arctic to northern temperate regions, and contains six species. The genus is characterized by the presence of the secondary metabolites pulvinic acid and vulpinic acid, compounds that when combined with usnic acid, give the species their characteristic yellow and green colors. Taxonomy The genus was circumscribed by Jan-Eric Mattson and Ming-Jou Lai in a 1993 ''Mycotaxon'' publication, to contain yellow species containing vulpinic and pinastric acids and a broadly club-shaped ascus. Mattson published a monograph of the genus later that year. The group of species assigned to the genus were previously recognized as a distinct grouping by Finnish lichenologist Veli Räsänen in 1952, who classified them in the genus ''Cetraria'', subgenus ''Platysma'', section ''Flavidae'', and subsection ''Cucullatae''. The t ...
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Ascocarp
An ascocarp, or ascoma (), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are most commonly bowl-shaped (apothecia) but may take on a spherical or flask-like form that has a pore opening to release spores (perithecia) or no opening (cleistothecia). Classification The ascocarp is classified according to its placement (in ways not fundamental to the basic taxonomy). It is called ''epigeous'' if it grows above ground, as with the morels, while underground ascocarps, such as truffles, are termed ''hypogeous''. The structure enclosing the hymenium is divided into the types described below (apothecium, cleistothecium, etc.) and this character ''is'' important for the taxonomic classification of the fungus. Apothecia can be relatively large and fleshy, whereas the others are microscopic—about the size of flecks ...
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Vulpinic Acid
Vulpinic acid is a natural product first found in and important in the symbiosis underlying the biology of lichens. It is a simple methyl ester derivative of its parent compound, pulvinic acid, and a close relative of pulvinone, both of which derive from aromatic amino acids such as phenylalanine via secondary metabolism. The roles of vulpinic acid are not fully established, but may include properties that make it an antifeedant for herbivores. The compound is relatively toxic to mammals. Chemical description Vulpinic acid was first isolated from lichens in 1925. As an isolated, purified substance, it is bright yellow in color. Vulpinic acid is derived biosynthetically by esterification from pulvinic acid; pulvinate itself derives from the aromatic amino acids phenylalanine and tyrosine, via dimerization and oxidative ring-cleavage of arylpyruvic acids, a process that also produces the related pulvinones. There have been several chemical syntheses reported for vulpinic acid. ...
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Vulpinic Acid
Vulpinic acid is a natural product first found in and important in the symbiosis underlying the biology of lichens. It is a simple methyl ester derivative of its parent compound, pulvinic acid, and a close relative of pulvinone, both of which derive from aromatic amino acids such as phenylalanine via secondary metabolism. The roles of vulpinic acid are not fully established, but may include properties that make it an antifeedant for herbivores. The compound is relatively toxic to mammals. Chemical description Vulpinic acid was first isolated from lichens in 1925. As an isolated, purified substance, it is bright yellow in color. Vulpinic acid is derived biosynthetically by esterification from pulvinic acid; pulvinate itself derives from the aromatic amino acids phenylalanine and tyrosine, via dimerization and oxidative ring-cleavage of arylpyruvic acids, a process that also produces the related pulvinones. There have been several chemical syntheses reported for vulpinic acid. ...
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Lichen Species
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (