Lessemsauridae
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Lessemsauridae
Lessemsauridae is a clade of early sauropodiform dinosaurs that lived in the Triassic and Jurassic of Argentina, South Africa and possibly Lesotho. A phylogenetic analysis performed by Apaldetti and colleagues in 2018 recovered a new clade of sauropodiforms uniting ''Lessemsaurus'', ''Antetonitrus'', and ''Ingentia'' which they named Lessemsauridae. It is a node-based taxon, defined as all descendants of the most recent common ancestor of ''Lessemsaurus sauropoides'' and ''Antetonitrus ingenipes''. Depending on the definition of Sauropoda, Lessemsauridae is either one of the most basal sauropod taxa, or a sister taxon of Sauropoda. An additional member of the clade was named later in 2018, ''Ledumahadi''. A 2021 study by Pol and colleagues also assigned the genera '' Kholumolumo'' and ''Meroktenos'' to the group. Lessemsaurids are recognised as very large quadrupeds that achieved giant sizes (up to 12 metric tons) independently of other giant sauropodomorphs. They had highly pne ...
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Ledumahadi
''Ledumahadi'' (meaning "giant thunderclap" in Sesotho language) is a genus of lessemsaurid sauropodomorph dinosaur from the Early Jurassic Elliot Formation in Free State Province, South Africa. The type and only species is ''L. mafube'', known from a singular incomplete postcranial specimen. A quadruped, it was one of the first giant sauropodomorphs, reaching a weight of around , despite not having evolved columnar limbs like its later huge relatives. Description ''Ledumahadi'' was likely a quadruped, as determined by analysis of the circumference of its humerus and femur compared to those of other dinosaurs. It would have had very large, robust forelimbs, consistent with those of its relatives. Unlike those of later sauropods, these limbs were naturally flexed, as opposed to being purely columnar. Size At its time in the Early Jurassic epoch, ''Ledumahadi'' is thought to have been the largest land animal that had ever lived. At the age of 14 years, ''L. mafube'' is estimated ...
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Lessemsaurus
''Lessemsaurus'' is an extinct genus of sauropod dinosaur belonging to Lessemsauridae. Naming and description The type species, ''L. sauropoides'', was formally described by José Fernando Bonaparte in 1999 in honor of Don Lessem, a writer of popular science books. It was found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in La Rioja Province, Argentina.Weishampel, David B; ''et al.'', 2004. "Dinosaur distribution (Late Triassic, South America)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 527–528. . This dinosaur was around long and was discovered in strata dating to the Norian stage, around 210 million years ago. It is estimated to have reached in maximum body mass. Classification A cladogram after Pol, Garrido & Cerda, 2011, illustrates a possible placing of ''Lessemsaurus'' and ''Antetonitrus'' in Sauropodomorpha: In 2018, Apaldetti ''et al.'' ...
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Meroktenos
''Meroktenos'' is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period of what is now Lesotho. Discovery and naming In 1959, François Ellenberger, Paul Ellenberger, Jean Fabre and Leonard Ginsburg discovered the type specimen, a thighbone or femur and other assorted bones, south of the village of Thabana Morena. In 1962 these were addressed in a thesis by D. Costedoat. The exact location the bones were recovered, is today unknown. In 1993, François-Xavier Gauffre assigned the remains to a second species of ''Melanorosaurus'': ''Melanorosaurus thabanensis''. The description was provisional, and in 1997 the fossil was described in more detail in a publication by Jacques van Heerden and Peter Malcolm Galton. The specific name refers to the site Thabana-Morena in Lesotho. Gauffre assumed that the specimen had been found in the Upper Elliot Formation dating from the Hettangian-Sinemurian and thus was about twenty million year younger than '' ...
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Antetonitrus
''Antetonitrus'' is a genus of sauropod dinosaur found in the Early Jurassic Elliot Formation of South Africa. The only species is ''Antetonitrus ingenipes''. As one of the oldest known sauropods, it is crucial for the understanding of the origin and early evolution of this group. It was a quadrupedal herbivore, like all of its later relatives, but shows primitive adaptations to use the forelimbs for grasping, instead of purely for weight support. Discovery and naming Adam Yates (paleontologist), Adam Yates, an Australian expert on early sauropodomorphs, named ''Antetonitrus'' in a 2003 report co-authored by South African James Kitching. The name is derived from the Latin ''ante-'' ("before") and ''tonitrus'' ("thunder"), which refers to its existence, before other known sauropods, specifically ''Brontosaurus'' ("thunder lizard"). The one known species of ''Antetonitrus'' is called ''A. ingenipes'', from the Latin ''ingens'' ("massive") and ''pes'' ("foot"), because it shows the ...
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Ingentia
''Ingentia'' is a genus of early sauropod dinosaur from the Late Triassic (late Norian-Rhaetian) of Argentina. The type specimen of ''Ingentia'', PVSJ 1086, was discovered in the Quebrada del Barro Formation of northwestern Argentina. It was described in 2018 by Cecilia Apaldetti, Ricardo Nestor Martínez, Ignacio Alejandro Cerda, Diego Pol and Oscar Alcober who named the type and only species ''Ingentia prima'', meaning "first huge one", as the taxon was one of the first very large sauropodomorphs to evolve, along with its close relative ''Lessemsaurus''. A second specimen, PVSJ 1087, was referred, containing five tail vertebrae, both ulnae and radii, a left calfbone and a right foot. Classification A phylogenetic analysis performed by Apaldetti ''et al.'' found a new clade uniting ''Ingentia'', ''Lessemsaurus'' and ''Antetonitrus'', which they named Lessemsauridae. Like other lessemsaurids, ''Ingentia'' has highly pneumatic and very antero-posteriorly short but tall robust cer ...
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Dinosaurs
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, are among ...
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Sauropoda
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their body), and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include ''Brachiosaurus'', ''Diplodocus'', ''Apatosaurus'' and ''Brontosaurus''. The oldest known unequivocal sauropod dinosaurs are known from the Early Jurassic. ''Isanosaurus'' and ''Antetonitrus'' were originally described as Triassic sauropods, but their age, and in the case of ''Antetonitrus'' also its sauropod status, were subsequently questioned. Sauropod-like sauropodomorph tracks from the Fleming Fjord Formation (Greenland) might, however, indicate the occurrence of the group in the Late Triassic. By the Late Jurassic (150 million yea ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Sister Group
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomic ...
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Metric Ton
The tonne ( or ; symbol: t) is a unit of mass equal to 1000 kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton to distinguish it from the non-metric units of the short ton (United States customary units), and the long ton ( British imperial units). It is equivalent to approximately 2204.6 pounds, 1.102 short tons, and 0.984 long tons. The official SI unit is the megagram (symbol: Mg), a less common way to express the same mass. Symbol and abbreviations The BIPM symbol for the tonne is t, adopted at the same time as the unit in 1879.Table 6
. BIPM. Retrieved on 2011-07-10.
Its use is also official for the metric ton in the United States, having been adopted by the United States

Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', t ...
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Cervical Vertebrae
In tetrapods, cervical vertebrae (singular: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In sauropsid species, the cervical vertebrae bear cervical ribs. In lizards and saurischian dinosaurs, the cervical ribs are large; in birds, they are small and completely fused to the vertebrae. The vertebral transverse processes of mammals are homologous to the cervical ribs of other amniotes. Most mammals have seven cervical vertebrae, with the only three known exceptions being the manatee with six, the two-toed sloth with five or six, and the three-toed sloth with nine. In humans, cervical vertebrae are the smallest of the true vertebrae and can be readily distinguished from those of the thoracic or lumbar regions by the presence of a foramen (hole) in each transverse process, through which the vertebral artery, vertebral veins, an ...
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