Ksar Metlili Formation
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Ksar Metlili Formation
The Ksar Metlili Formation is a geological formation in eastern High Atlas of Morocco, it is late Tithonian to Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago) ... in age. It is approximately thick and primarily consists of mudstone and sandstone, with thin calcareous beds. One of these calcareous beds near the middle of the sequence is an important microvertebrate locality. Subsequent to the original site, several other localities have been sampled. The depositional environment is thought to be near shore deltaic. Fossil content Amphibians Lepidosaurs Dinosaurs Mammals References {{reflist Geologic formations of Morocco Jurassic System of Africa Tithonian Stage Jurassic Morocco Upper Cretaceous Series of Africa Berriasian Stage Cretaceous Mor ...
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Tithonian
In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the Kimmeridgian and followed by the Berriasian (part of the Cretaceous).See for a detailed version of the geologic timescale Gradstein ''et al.'' (2004) Stratigraphic definitions The Tithonian was introduced in scientific literature by German stratigrapher Albert Oppel in 1865. The name Tithonian is unusual in geological stage names because it is derived from Greek mythology. Tithonus was the son of Laomedon of Troy and fell in love with Eos, the Greek goddess of dawn. His name was chosen by Albert Oppel for this stratigraphical stage because the Tithonian finds itself hand in hand with the dawn of the Cretaceous. The base of the Tithonian stage is at the base of the ammonite biozone of '' Hybonoticeras hybonotum''. A global reference profi ...
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Theropoda
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic Period (geology), period 231.4 million years ago (Year#SI prefix multipliers, Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of di ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, are among ...
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Rhynchocephalia
Rhynchocephalia (; ) is an order of lizard-like reptiles that includes only one living species, the tuatara (''Sphenodon punctatus'') of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a diverse group including a wide array of morphologically distinct forms. The oldest record of the group is dated to the Middle Triassic around 238 to 240 million years ago, and they had achieved a worldwide distribution by the Early Jurassic. Most rhynchocephalians belong to the group Sphenodontia ('wedge-teeth'). Their closest living relatives are lizards and snakes in the order Squamata, with the two orders being grouped together in the superorder Lepidosauria. Many of the niches occupied by lizards today were held by sphenodontians during the Triassic and Jurassic, although lizard diversity began to overtake sphenodontian diversity in the Cretaceous, and they had disappeared almost entirely by the beginning of the Cenozoic. While the modern tuat ...
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Tingitana Anoualae
Mauretania Tingitana (Latin for "Tangerine Mauretania") was a Roman province, coinciding roughly with the northern part of present-day Morocco. The territory stretched from the northern peninsula opposite Gibraltar, to Sala Colonia (or Chellah) and Volubilis to the south, and as far east as the Mulucha (or Malva) river. Its capital city was Tingis, which is the modern Tangier. Other major cities of the province were Iulia Valentia Banasa, Septem, Rusadir, Lixus and Tamuda. History After the death in 40 AD of Ptolemy of Mauretania, the last Ptolemaic ruler of the Kingdom of Mauretania, in about 44 AD Roman Emperor Claudius annexed the kingdom to the Roman Empire and partitioned it into two Roman provinces: Mauretania Tingitana and Mauretania Caesariensis. The Mulucha (Moulouya River), located around 60 km west of modern Oran, Algeria, became the border separating them. The Roman occupation did not extend very far into the continent. In the far west, the southern limit o ...
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Paramacellodus
''Paramacellodus'' is an extinct genus of scincomorph lizards from the Early Cretaceous of England and France, and the Late Jurassic of Portugal and the western United States. The type species, ''Paramacellodus oweni'', was named in 1967 from the earliest Cretaceous (Berriasian) Purbeck Group in Dorset, England. Additional material referable to a species of ''Paramacellodus'', possibly ''P. oweni'', has been described from the Morrison Formation, specifically in Como Bluff, Wyoming, and Dinosaur National Monument, Utah.Foster, J. (2007). "Table 2.1: Fossil Vertebrates of the Morrison Formation." ''Jurassic West: The Dinosaurs of the Morrison Formation and Their World''. Indiana University Press. pp. 58–59. An indeterminate species is known from the Berriasian aged Angeac-Charente bonebed in France.Ronan Allain, Romain Vullo, Lee Rozada, Jérémy Anquetin, Renaud Bourgeais, et al.Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southw ...
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Lepidosauria
The Lepidosauria (, from Greek meaning ''scaled lizards'') is a subclass or superorder of reptiles, containing the orders Squamata and Rhynchocephalia. Squamata includes snakes, lizards, and amphisbaenians. Squamata contains over 9,000 species, making it by far the most species-rich and diverse order of reptiles in the present day. Rhynchocephalia was a formerly widespread and diverse group of reptiles in the Mesozoic Era. However, it is represented by only one living species: the tuatara (''Sphenodon punctatus),'' a superficially lizard-like reptile native to New Zealand. Lepidosauria is a monophyletic group (i.e. a clade), containing all descendants of the last common ancestor of squamates and rhynchocephalians. Lepidosaurs can be distinguished from other reptiles via several traits, such as large keratinous scales which may overlap one another. Purely in the context of modern taxa, Lepidosauria can be considered the sister taxon to Archosauria, which includes Aves (birds) and ...
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Caecilian
Caecilians (; ) are a group of limbless, vermiform or serpentine amphibians. They mostly live hidden in the ground and in stream substrates, making them the least familiar order of amphibians. Caecilians are mostly distributed in the tropics of South and Central America, Africa, and southern Asia. Their diet consists of small subterranean creatures such as earthworms. All modern caecilians and their closest fossil relatives are grouped as a clade, Apoda , within the larger group Gymnophiona , which also includes more primitive extinct caecilian-like amphibians. The name derives from the Greek words γυμνος (''gymnos'', naked) and οφις (''ophis'', snake), as the caecilians were originally thought to be related to snakes. The body is cylindrical dark brown or bluish black in colour. The skin is slimy and bears grooves or ringlike markings. Description Caecilians completely lack limbs, making the smaller species resemble worms, while the larger species, with lengths up ...
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Rubricacaecilia
''Rubricacaecilia'' is an extinct genus of caecilian (limbless amphibian) from the Berriasian aged Ksar Metlili Formation in Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ....Evans, S. E. & Sigogneau-Rusell, D. (2001A stem-group caecilian (Amphibia: Lissamphibia) from the Lower Cretaceous of Morocco ''Palaeontology'' 44, 259–273. References Gymnophiona Early Cretaceous amphibians of Africa Fossil taxa described in 2001 Prehistoric amphibian genera Berriasian genera {{paleo-amphibian-stub ...
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