Karaurus
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Karaurus
''Karaurus'' (meaning ''head-tail'') is an extinct genus of stem-group salamander (Caudata) from the Middle to Late Jurassic (Callovian–Kimmeridgian) Karabastau Formation of Kazakhstan. It is one of the oldest salamanders known. ''Karaurus'' was large for a Jurassic salamander, about long, and very similar anatomically to modern salamanders. ''Karaurus'' is thought to have fed using suction feeding via the enlargement of the buccal cavity on small fish and invertebrates, with the well developed palatal dentition (teeth on the roof of the mouth) and marginal teeth helping to grasp prey. ''Karaurus'' is thought to form a clade with ''Kokartus'' from the Middle Jurassic (Bathonian) of Kyrgyzstan, and ''Marmorerpeton'' from the Bathonian of Britain, together forming the Karauridae, which are closely related to crown salamanders. Like other members of Karauridae, ''Karaurus'' is neotenic Neoteny (), also called juvenilization,Montagu, A. (1989). Growing Young. Bergin & Garvey: CT ...
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Salamander
Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela. Salamander diversity is highest in eastern North America, especially in the Appalachian Mountains; most species are found in the Holarctic realm, with some species present in the Neotropical realm. Salamanders rarely have more than four toes on their front legs and five on their rear legs, but some species have fewer digits and others lack hind limbs. Their permeable skin usually makes them reliant on habitats in or near water or other cool, damp places. Some salamander species are fully aquatic throughout their lives, some take to the water intermittently, and others are entirely terrestrial as adults. This group of amphibians is capable of regenerating lost lim ...
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Karauridae
The Karauridae are a family of stem-group salamanders (Caudata) that are known from the Middle Jurassic to Late Jurassic in Central Asia (Kazakhstan and Kyrgyzstan) and Western Europe (United Kingdom). The family includes three members: ''Karaurus'' from the Middle-Late Jurassic Karabastau Formation of Kazakhstan, ''Kokartus'' from the Middle Jurassic Balabansai Formation of Kyrgyzstan, and ''Marmorerpeton'' from the Middle Jurassic Forest Marble Formation of England and Kilmaluag Formation of Scotland. The members are some of the oldest known salamanders. The family is united by several morphological characters, including sculptured skull roof bones. Like some modern salamanders, karaurids were neotenic. Members of the family likely fed via suction feeding Aquatic feeding mechanisms face a special difficulty as compared to feeding on land, because the density of water is about the same as that of the prey, so the prey tends to be pushed away when the mouth is closed. This probl ...
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Caudata
The Caudata are a group of amphibians containing the extant salamanders (Urodela) and all extinct species of amphibians more closely related to salamanders than to frogs. They are typically characterized by a superficially lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. Disagreement exists between different authorities as to the definition of the terms "Caudata" and "Urodela". Some maintain that Urodela should be restricted to the crown group, with Caudata being used for the total group. Others restrict the name Caudata to the crown group and use Urodela for the total group. The former approach seems to be most widely adopted and is used in this article. Evolution The origins and evolutionary relationships between the three main groups of amphibians ( apodans, urodeles and anurans) is a matter of debate. A 2005 molecular phylogeny, based on rDNA analysis, suggest ...
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Karabastau Formation
The Karabastau Formation ( kk, Qarabastaý svıtasy) is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that has been studied since the early 20th century, alongside the rarer remains of vertebrates, including pterosaurs, salamanders, lizards and crocodiles.Barrett, P.M., Butler, R.J., Edwards, N.P., & Milner, A.R. Pterosaur distribution in time and space: an atlas. p61-107. in Flugsaurier: Pterosaur papers in honour of Peter Wellnhofer. 2008. Hone, D.W.E., and Buffetaut, E. (eds). Zitteliana B, 28. 264p/ref> Lithology and depositional environment The primary lithology consists of 1 mm thick varve laminations of claystone, with a dark part and a light dolomitic part, which probably correspond to a wet and dry season respectively, alongside rare, several cm thick sandstone interbeds. These were deposited within an ancient freshwater paleol ...
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Marmorerpeton
''Marmorerpeton'' is an extinct genus of prehistoric stem group-salamanders that lived in Britain during the Bathonian stage of the Middle Jurassic. They are among the oldest known salamanders. Two species were named when the genus was first described by Susan E. Evans et al. in 1988, ''M''. ''freemani'', and ''M''. ''kermacki'', from fragmentary remains found via screenwashing in the Forest Marble Formation of England. Due to the size of their osteocytic lacunae suggesting a large genome size and some morphological characters, like the presence of calcified cartilage in the medulla of its humerus, it was assumed that ''Marmorerpeton'' was neotenic. New more complete remains of a new species ''M. wakei'' were described in 2022 from the Kilmaluag Formation of the Isle of Skye, Scotland. These conclusively demonstrated that ''Marmorerpeton'' was neotenic, and was a member of the family Karauridae, with the other two members of the family, ''Karaurus'' and ''Kokartus'' being known ...
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Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name "Malm" indicates rocks of Late Jurassic age. In the past, ''Malm'' was also used to indicate the unit of geological time, but this usage is now discouraged to make a clear distinction between lithostratigraphic and geochronologic/chronostratigraphic units. Subdivisions The Late Jurassic is divided into three ages, which correspond with the three (faunal) stages of Upper Jurassic rock: Paleogeography During the Late Jurassic Epoch, Pangaea broke up into two supercontinents, Laurasia to the north, and Gondwana to the south. The result of this break-up was the spawning of the Atlantic Ocean. However, at this time, the Atlantic Ocean was relatively narrow. Life forms of the epoch This epoch is well known for many famous types of dinosau ...
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Prehistoric Amphibians Of Asia
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. ...
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Late Jurassic Amphibians
Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, a concept in econometrics Music * ''Late'' (album), a 2000 album by The 77s * Late!, a pseudonym used by Dave Grohl on his ''Pocketwatch'' album * Late (rapper), an underground rapper from Wolverhampton * "Late" (song), a song by Blue Angel * "Late", a song by Kanye West from ''Late Registration'' Other * Late (Tonga), an uninhabited volcanic island southwest of Vavau in the kingdom of Tonga * "Late" (''The Handmaid's Tale''), a television episode * LaTe, Oy Laivateollisuus Ab, a defunct shipbuilding company * Late may refer to a person who is Dead See also * * * ''Lates'', a genus of fish in the lates perch family * Later (other) * Tardiness * Tardiness (scheduling) In scheduling, tardiness is a measure of a delay in exe ...
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Prehistoric Salamanders
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. ...
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Neotenic
Neoteny (), also called juvenilization,Montagu, A. (1989). Growing Young. Bergin & Garvey: CT. is the delaying or slowing of the physiological, or somatic, development of an organism, typically an animal. Neoteny is found in modern humans compared to other primates. In progenesis or paedogenesis, sexual development is accelerated. Both neoteny and progenesis result in paedomorphism (as having the form typical of children) or paedomorphosis (changing towards forms typical of children), a type of heterochrony. It is the retention in adults of traits previously seen only in the young. Such retention is important in evolutionary biology, domestication and evolutionary developmental biology. Some authors define paedomorphism as the retention of larval traits, as seen in salamanders.Schell, S. C. ''Handbook of Trematodes of North America North of Mexico'', 1985, pg. 22 History and etymology The origins of the concept of neoteny have been traced to the Bible (as argued by Ashley Monta ...
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Buccal Cavity
The buccal space (also termed the buccinator space) is a fascial spaces of the head and neck, fascial space of the head and neck (sometimes also termed fascial tissue spaces or tissue spaces). It is a potential space in the cheek, and is paired on each side. The buccal space is superficial to the buccinator muscle and deep to the platysma muscle and the skin. The buccal space is part of the Subcutaneous tissue, subcutaneous space, which is continuous from head to toe. Structure Boundaries The boundaries of each buccal space are: * the angle of the mouth anteriorly, * the masseter muscle posteriorly, * the zygomatic process of the maxilla and the zygomaticus muscles superiorly, * the depressor anguli oris muscle and the attachment of the deep fascia to the mandible inferiorly, * the buccinator muscle medially (the buccal space is superficial to the buccinator), * the platysma muscle, subcutaneous tissue and skin laterally (the space is deep to platysma). Communications * to the pte ...
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Bathonian
In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Stratigraphic definitions The Bathonian Stage takes its name from Bath, a spa town in England built on Jurassic limestone (the Latinized form of the town name is ''Bathonium''). The name was introduced in scientific literature by Belgian geologist d'Omalius d'Halloy in 1843. The original type locality was located near Bath. The French palaeontologist Alcide d'Orbigny was in 1852 the first to define the exact length of the stage. The base of the Bathonian is at the first appearance of ammonite species '' Parkinsonia (Gonolkites) convergens'' in the stratigraphic column. The global reference profile for the base of the Bathonian (a GSSP) was ratified as Ravin du Bès, Bas-Auran area, Alpes de Haute Provence, France in 2009. The top of the Bat ...
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