Isoëtales
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Isoëtales
Isoetales, sometimes also written Isoëtales, is an order of plants in the class Lycopodiopsida. There are about 140-150 living species, all of which are classified in the genus '' Isoetes'' (quillworts), with a cosmopolitan distribution, but often scarce to rare. Living species are mostly aquatic or semi-aquatic, and are found in clear ponds and slowly moving streams. Each leaf is slender and broadens downward to a swollen base up to 5 mm wide where the leaves attach in clusters to a bulb-like, underground corm characteristic of most quillworts. This swollen base also contains male and female sporangia, protected by a thin, transparent covering ( velum), which is used diagnostically to help identify quillwort species. Quillwort species are very difficult to distinguish by general appearance. The best way to identify them is by examining the megaspores under a microscope. '' Isoetes'' are the only living pteridophytes capable of secondary growth. Fossils Fossilised ...
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Secondary Growth
In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of stems and roots, causing them to elongate, and gives rise to primary tissue. Secondary growth occurs in most seed plants, but monocots usually lack secondary growth. If they do have secondary growth, it differs from the typical pattern of other seed plants. The formation of secondary vascular tissues from the cambium is a characteristic feature of dicotyledons and gymnosperms. In certain monocots, the vascular tissues are also increased after the primary growth is completed but the cambium of these plants is of a different nature. In the living pteridophytes this feature is extremely rare, only occurring in ''Isoetes''. Lateral meristems In many vascular plants, secondary growth is the result of the activity of the two lateral meriste ...
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Pleuromeia
''Pleuromeia'' is an extinct genus of lycophytes. They are related to modern quillworts (''Isoetes''). ''Pleuromeia'' dominated vegetation during the Early Triassic all over Eurasia and elsewhere, in the aftermath of the collapse of floral communities during the Permian–Triassic extinction event, often occurring in monospecific assemblages. Its sedimentary context in monospecific assemblages on immature paleosols, is evidence that it was an opportunistic pioneer plant that grew on mineral soils with little competition. It spread to high latitudes with greenhouse climatic conditions. Conifers reoccurred in the Early Anisian, followed by the cycads and pteridosperms during the Late Anisian. Description ''Pleuromeia'' consisted of a single unbranched stem of variable thickness, which could grow to a maximum of in height. Around the stem were helically arranged leaves. It had a 2-4 lobed bulbous base to which numerous adventive roots are attached. ''Pleuromeia'' produced a s ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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Spore
In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. Under favourable conditions the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote which develops into a new s ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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Isoetes Beestonii
''Isoetes beestonii'' is the oldest known species of the living quillwort genus from the latest Permian of New South Wales and Queensland. Originally considered earliest Triassic, it is now known to be latest Permian in age, immediately before the Permian Triassic mass extinction. Description ''Isoetes beestonii'' is preserved as whole plants in life position within bedding planes, and presumably lived as an early successional weed in lake and pond sedimentary environments, like living ''Isoetes''. Its leaves were wider and more succulent than modern species of ''Isoetes''. Like modern ''Isoetes'', fertile plants were little different from sterile plants, unlike Early Triassic ''Tomiostrobus'' which formed woody conelike fertile plants. See also * Evolution of plants The evolution of plants has resulted in a wide range of complexity, from the earliest algal mats, through multicellular marine and freshwater green algae, terrestrial bryophytes, lycopods and ferns, to the ...
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Pleuromeia Restoration
''Pleuromeia'' is an extinct genus of lycophytes. They are related to modern quillworts (''Isoetes''). ''Pleuromeia'' dominated vegetation during the Early Triassic all over Eurasia and elsewhere, in the aftermath of the collapse of floral communities during the Permian–Triassic extinction event, often occurring in monospecific assemblages. Its sedimentary context in monospecific assemblages on immature paleosols, is evidence that it was an opportunistic pioneer plant that grew on mineral soils with little competition. It spread to high latitudes with greenhouse climatic conditions. Conifers reoccurred in the Early Anisian, followed by the cycads and pteridosperms during the Late Anisian. Description ''Pleuromeia'' consisted of a single unbranched stem of variable thickness, which could grow to a maximum of in height. Around the stem were helically arranged leaves. It had a 2-4 lobed bulbous base to which numerous adventive roots are attached. ''Pleuromeia'' produced ...
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Pteridophyte
A pteridophyte is a vascular plant (with xylem and phloem) that disperses spores. Because pteridophytes produce neither flowers nor seeds, they are sometimes referred to as "cryptogams", meaning that their means of reproduction is hidden. Ferns, horsetails (often treated as ferns), and lycophytes (clubmosses, spikemosses, and quillworts) are all pteridophytes. However, they do not form a monophyletic group because ferns (and horsetails) are more closely related to seed plants than to lycophytes. "Pteridophyta" is thus no longer a widely accepted taxon, but the term ''pteridophyte'' remains in common parlance, as do ''pteridology'' and ''pteridologist'' as a science and its practitioner, respectively. Ferns and lycophytes share a life cycle and are often collectively treated or studied, for example by the International Association of Pteridologists and the Pteridophyte Phylogeny Group. Description Pteridophytes (ferns and lycophytes) are free-sporing vascular plants that have a lif ...
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