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Inclusive Fitness
In evolutionary biology, inclusive fitness is one of two metrics of evolutionary success as defined by W. D. Hamilton in 1964: * Personal fitness is the number of offspring that an individual begets (regardless of who rescues/rears/supports them) * Inclusive fitness is the number of offspring equivalents that an individual rears, rescues or otherwise supports through its behaviour (regardless of who begets them) An individual's own child, who carries one half of the individual's genes, is defined as one offspring equivalent. A sibling's child, who will carry one-quarter of the individual's genes, is 1/2 offspring equivalent. Similarly, a cousin's child, who has 1/16 of the individual's genes, is 1/8 offspring equivalent. From the gene's point of view, evolutionary success ultimately depends on leaving behind the maximum number of copies of itself in the population. Prior to Hamilton's work, it was generally assumed that genes only achieved this through the number of viable o ...
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Evolutionary Biology
Evolutionary biology is the subfield of biology that studies the evolutionary processes (natural selection, common descent, speciation) that produced the diversity of life on Earth. It is also defined as the study of the history of life forms on Earth. Evolution is based on the theory that all species are related and they gradually change over time. In a population, the genetic variations affect the physical characteristics i.e. phenotypes of an organism. These changes in the phenotypes will be an advantage to some organisms, which will then be passed onto their offspring. Some examples of evolution in species over many generations are the Peppered Moth and Flightless birds. In the 1930s, the discipline of evolutionary biology emerged through what Julian Huxley called the modern synthesis of understanding, from previously unrelated fields of biological research, such as genetics and ecology, systematics, and paleontology. The importance of studying Evolutionary bio ...
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Green-beard Effect
The green-beard effect is a thought experiment used in evolutionary biology to explain selective altruism among individuals of a species. The idea of a green-beard gene was proposed by William D. Hamilton in his articles of 1964, and got the name from the example used by Richard Dawkins (''"I have a green beard and I will be altruistic to anyone else with green beard"'') in '' The Selfish Gene'' (1976). A green-beard effect occurs when an allele, or a set of linked alleles, produce three expressed (or phenotypic) effects: * a perceptible trait—the hypothetical "green beard" * recognition of this trait by others; and * preferential treatment of individuals with the trait by others with the trait The carrier of the gene (or a specific allele) is essentially recognizing copies of the same gene (or a specific allele) in other individuals. Whereas kin selection involves altruism to related individuals who share genes in a non-specific way, green-beard alleles promote al ...
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Robin Fox
Robin Fox (born 1934) is an Anglo-American anthropologist who has written on the topics of incest avoidance, marriage systems, human and primate kinship systems, evolutionary anthropology, sociology and the history of ideas in the social sciences. He founded the department of anthropology at Rutgers University in 1967 and had remained a professor there for the rest of his career, also being a director of research for the H. F. Guggenheim Foundation from 1972 to 1984. Fox published ''The Imperial Animal'', with Lionel Tiger in 1971 one of the earliest to advocate and demonstrate an evolutionary approach to the understanding of human social behaviour. His daughter Kate Fox wrote the book ''Watching the English.'' In 2013 Fox was elected to the US National Academy of Sciences (Anthropology and Evolutionary Biology). Life and work Robin Fox was born in the village of Haworth in the Yorkshire Dales, at the nadir of the Great Depression in 1934. He had very little schooling duri ...
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R/K Selection Theory
In ecology, ''r''/''K'' selection theory relates to the selection of combinations of traits in an organism that trade off between quantity and quality of offspring. The focus on either an increased quantity of offspring at the expense of individual parental investment of ''r''-strategists, or on a reduced quantity of offspring with a corresponding increased parental investment of ''K''-strategists, varies widely, seemingly to promote success in particular environments. The concepts of quantity or quality offspring are sometimes referred to as "cheap" or "expensive", a comment on the expendable nature of the offspring and parental commitment made. The stability of the environment can predict if many expendable offspring are made or if fewer offspring of higher quality would lead to higher reproductive success. An unstable environment would encourage the parent to make many offspring, because the likelihood of all (or the majority) of them surviving to adulthood is slim. In contra ...
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Reproductive Success
Reproductive success is an individual's production of offspring per breeding event or lifetime. This is not limited by the number of offspring produced by one individual, but also the reproductive success of these offspring themselves. Reproductive success is different from fitness in that individual success is not necessarily a determinant for adaptive strength of a genotype since the effects of chance and the environment have no influence on those specific genes. Reproductive success turns into a part of fitness when the offspring are actually recruited into the breeding population. If offspring quantity is not correlated with quality this holds up, but if not then reproductive success must be adjusted by traits that predict juvenile survival in order to be measured effectively. Quality and quantity is about finding the right balance between reproduction and maintenance. The disposable soma theory of aging tells us that a longer lifespan will come at the cost of reproduction ...
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Kin Selection
Kin selection is the evolutionary strategy that favours the reproductive success of an organism's relatives, even when at a cost to the organism's own survival and reproduction. Kin altruism can look like altruistic behaviour whose evolution is driven by kin selection. Kin selection is an instance of inclusive fitness, which combines the number of offspring produced with the number an individual can ensure the production of by supporting others, such as siblings. Charles Darwin discussed the concept of kin selection in his 1859 book, ''On the Origin of Species'', where he reflected on the puzzle of sterile social insects, such as honey bees, which leave reproduction to their mothers, arguing that a selection benefit to related organisms (the same "stock") would allow the evolution of a trait that confers the benefit but destroys an individual at the same time. R.A. Fisher in 1930 and J.B.S. Haldane in 1932 set out the mathematics of kin selection, with Haldane famously joki ...
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Hamiltonian Spite
Within the field of social evolution, Hamiltonian spite is a term for spiteful behaviours occurring among conspecifics that have a cost for the actor and a negative impact upon the recipient. Theories on altruism and spitefulness W. D. Hamilton published an influential paper on altruism in 1964 to explain why genetic kin tend to help each other. He argued that genetically related individuals are likely to carry the copies of the same alleles; thus, helping kin may ensure that copies of the actors' alleles pass onto next generations of both the recipient and the actor. While this became a widely accepted idea, it was less noted that Hamilton published a later paper that modified this view. This paper argues that by measuring the genetic relatedness between any two (randomly chosen) individuals of a population several times, we can identify an average level of relatedness. Theoretical models predict that (1) it is adaptive for an individual to be altruistic to any other indivi ...
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Evolutionary Psychology
Evolutionary psychology is a theoretical approach in psychology that examines cognition and behavior from a modern evolutionary perspective. It seeks to identify human psychological adaptations with regards to the ancestral problems they evolved to solve. In this framework, psychological traits and mechanisms are either functional products of natural and sexual selection, non-adaptive by-products of other adaptive traits, or noise. Adaptationist thinking about physiological mechanisms, such as the heart, lungs, and the liver, is common in evolutionary biology. Evolutionary psychologists apply the same thinking in psychology, arguing that just as the heart evolved to pump blood, and the liver evolved to detoxify poisons, there is modularity of mind in that different psychological mechanisms evolved to solve different adaptive problems. These evolutionary psychologists argue that much of human behavior is the output of psychological adaptations that evolved to solve recurr ...
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Criticism Of Evolutionary Psychology
Evolutionary psychology seeks to identify and understand human psychological traits that have evolved in much the same way as biological traits, through adaptation to environmental cues. Furthermore, it tends toward viewing the vast majority of psychological traits, certainly the most important ones, as the result of past adaptions, which has generated significant controversy and criticism from competing fields. These criticisms include disputes about the testability of evolutionary hypotheses, cognitive assumptions such as massive modularity, vagueness stemming from assumptions about the environment that leads to evolutionary adaptation, the importance of non-genetic and non-adaptive explanations, as well as political and ethical issues in the field itself. Evolutionary psychologists contend the criticisms against it are straw men, based on an incorrect nature versus nurture dichotomy, and/or based on misunderstandings of the discipline.Alcock, John (2001). The Triumph of Sociobio ...
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The New York Times
''The New York Times'' (''the Times'', ''NYT'', or the Gray Lady) is a daily newspaper based in New York City with a worldwide readership reported in 2020 to comprise a declining 840,000 paid print subscribers, and a growing 6 million paid digital subscribers. It also is a producer of popular podcasts such as '' The Daily''. Founded in 1851 by Henry Jarvis Raymond and George Jones, it was initially published by Raymond, Jones & Company. The ''Times'' has won 132 Pulitzer Prizes, the most of any newspaper, and has long been regarded as a national " newspaper of record". For print it is ranked 18th in the world by circulation and 3rd in the U.S. The paper is owned by the New York Times Company, which is publicly traded. It has been governed by the Sulzberger family since 1896, through a dual-class share structure after its shares became publicly traded. A. G. Sulzberger, the paper's publisher and the company's chairman, is the fifth generation of the family to head the ...
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Ad Hoc
Ad hoc is a Latin phrase meaning literally 'to this'. In English, it typically signifies a solution for a specific purpose, problem, or task rather than a generalized solution adaptable to collateral instances. (Compare with ''a priori''.) Common examples are ad hoc committees and commissions created at the national or international level for a specific task. In other fields, the term could refer to, for example, a military unit created under special circumstances (see '' task force''), a handcrafted network protocol (e.g., ad hoc network), a temporary banding together of geographically-linked franchise locations (of a given national brand) to issue advertising coupons, or a purpose-specific equation. Ad hoc can also be an adjective describing the temporary, provisional, or improvised methods to deal with a particular problem, the tendency of which has given rise to the noun ''adhocism''. Styling Style guides disagree on whether Latin phrases like ad hoc should be italicize ...
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Martin Nowak
Martin Andreas Nowak (born April 7, 1965) is an Austrian-born professor of mathematical biology, at Harvard University since 2003. He is one of the leading researchers in the field that studies the role of cooperation in evolution. Nowak has held professorships in Oxford and Princeton before being recruited to Harvard in 2003 when Jeffrey Epstein donated a large sum of money to set up a center for studying cooperation in evolution. Nowak's best known work outside of academia is his 2011 book ''SuperCooperators: Altruism, Evolution and Why We Need Each Other to Succeed''. This book is partly an autobiography and partly a popular presentation of his work in mathematical biology on the evolution of cooperation. In the book Nowak also wrote favorably about his interactions with Jeffrey Epstein, who had donated to Harvard to enable the creation of an institute headed by Nowak. In 2020 Nowak was disciplined by being suspended from teaching for two years and having his institute perma ...
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