Hornwort
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Hornwort
Hornworts are a group of non-vascular Embryophytes (land plants) constituting the division Anthocerotophyta (). The common name refers to the elongated horn-like structure, which is the sporophyte. As in mosses and liverworts, hornworts have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of genetic information; the flattened, green plant body of a hornwort is the gametophyte stage of the plant. Hornworts may be found worldwide, though they tend to grow only in places that are damp or humid. Some species grow in large numbers as tiny weeds in the soil of gardens and cultivated fields. Large tropical and sub-tropical species of ''Dendroceros'' may be found growing on the bark of trees. The total number of species is still uncertain. While there are more than 300 published species names, the actual number could be as low as 100-150 species. Description Like all bryophytes, the dominant life phase of a hornwort is the haploid gametophyte. ...
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Plant
Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclude the fungi and some algae, as well as the prokaryotes (the archaea and bacteria). By one definition, plants form the clade Viridiplantae (Latin name for "green plants") which is sister of the Glaucophyta, and consists of the green algae and Embryophyta (land plants). The latter includes the flowering plants, conifers and other gymnosperms, ferns and their allies, hornworts, liverworts, and mosses. Most plants are multicellular organisms. Green plants obtain most of their energy from sunlight via photosynthesis by primary chloroplasts that are derived from endosymbiosis with cyanobacteria. Their chloroplasts contain chlorophylls a and b, which gives them their green color. Some plants are parasitic or mycotrophic and have lost the ...
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Embryophytes
The Embryophyta (), or land plants, are the most familiar group of green plants that comprise vegetation on Earth. Embryophytes () have a common ancestor with green algae, having emerged within the Phragmoplastophyta clade of green algae as sister of the Zygnematophyceae. The Embryophyta consist of the bryophytes plus the polysporangiophytes. Living embryophytes therefore include hornworts, liverworts, mosses, lycophytes, ferns, gymnosperms and flowering plants. The land plants have diplobiontic life cycles and it is accepted now that they emerged from freshwater, multi-celled algae. The embryophytes are informally called land plants because they live primarily in terrestrial habitats (with exceptional members who evolved to live once again in aquatic habitats), while the related green algae are primarily aquatic. Embryophytes are complex multicellular eukaryotes with specialized reproductive organs. The name derives from their innovative characteristic of nurturing the young ...
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Dendroceros
''Dendroceros'' is a genus of hornworts in the family Dendrocerotaceae.Renzaglia, Karen S. & Kevin C. Vaughn. (2000) "Anatomy, development and classification of hornworts", pages 1-20 ''in'' A. Jonathan Shaw & Bernard Goffinet (Eds.), ''Bryophyte Biology''. (Cambridge: Cambridge University Press). The genus contains about 51 species native to tropical and sub-tropical regions of the world. Description The epiphytic and epiphyllous Dendroceros is the only desiccation-tolerant hornwort genus. The gametophyte is yellowish-green and usually less than one-half cm wide. The thallus branches in a bifurcating pattern. In the subgenus ''Apoceros'', there are cavities in the central strand of the thallus. The edges of the thallus are only a single layer of cells thick and have an undulating margin. It is common to find symbiotic colonies of blue-green bacteria (usually ''Nostoc'') growing among the cells. Under a microscope, the epidermal cells have trigones. The sporophyte is erect w ...
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Leiosporoceros
''Leiosporoceros dussii'' is the only species in the hornwort genus ''Leiosporoceros''. The species is placed in a separate family, order, and class for being " genetically and morphologically distinct from all other hornwort lineages." Cladistic analysis of genetic data supports a position at the very base of the hornwort clade. Physical characteristics that distinguish the group include unusually small spores that are ''monolete'' and unornamented. Additionally, there are unique strands of ''Nostoc'' (cyanobacteria) that grow inside the plant parallel with its direction of growth. Unlike other hornworts with symbiotic cyanobacteria that enters through mucilage clefts, the mucilage clefts in Leiosporoceros is only present in young plants and then closes permanently once the cyanobacterial colonies have been established. Also mycorrhiza and pyrenoids are absent. Male plants have been found in Panama. The Ordovician fossil A fossil (from Classical Latin , ) is any pre ...
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Anthocerotaceae
The Anthocerotaceae is the only family of hornwort Hornworts are a group of non-vascular Embryophytes (land plants) constituting the division Anthocerotophyta (). The common name refers to the elongated horn-like structure, which is the sporophyte. As in mosses and liverworts, hornworts have a ...s in the order Anthocerotales. References External links Hornworts Hornwort families {{Bryophyte-stub ...
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Dendrocerotaceae
The Dendrocerotaceae is the only family of hornwort Hornworts are a group of non-vascular Embryophytes (land plants) constituting the division Anthocerotophyta (). The common name refers to the elongated horn-like structure, which is the sporophyte. As in mosses and liverworts, hornworts have a ...s in the order Dendrocerotales. Phylogeny Currently phylogeny. References External links Hornworts Hornwort families {{Bryophyte-stub ...
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Notothyladaceae
The Notothyladaceae is the only family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ... of hornworts in the Order (biology), order Notothyladales. In Guizhou Province, SW China, a new species named Notothylas guizhouensis is reported (ZHANG 2018). The species is easily distinguished from congeners by: the lack of a columella, the dehisence line consisting of two rows of brown, thick-walled cells, the absence of lamellae in the involucre, the epidermal cells of the capsule having a moderately thick wall, the lack of an easily recognisable equatorial girdle of the spore, and mature spores dark brown; the proximal (ZHANG 2018). ZHANG, ZUO, Q., LI, J., & PENG, T. (2018). A new species of Notothylas (Notothyladaceae) from southwest China. Phytotaxa, 367(2), 191–. https://doi.org/10 ...
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Liverworts
The Marchantiophyta () are a division of non-vascular land plants commonly referred to as hepatics or liverworts. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of genetic information. It is estimated that there are about 9000 species of liverworts. Some of the more familiar species grow as a flattened leafless thallus, but most species are leafy with a form very much like a flattened moss. Leafy species can be distinguished from the apparently similar mosses on the basis of a number of features, including their single-celled rhizoids. Leafy liverworts also differ from most (but not all) mosses in that their leaves never have a costa (present in many mosses) and may bear marginal cilia (very rare in mosses). Other differences are not universal for all mosses and liverworts, but the occurrence of leaves arranged in three ranks, the presence of deep lobes or segmented leaves, or a lack of clearly differ ...
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Pyrenoid
Pyrenoids are sub-cellular micro-compartments found in chloroplasts of many algae,Giordano, M., Beardall, J., & Raven, J. A. (2005). CO2 concentrating mechanisms in algae: mechanisms, environmental modulation, and evolution. Annu. Rev. Plant Biol., 56, 99-131. and in a single group of land plants, the hornworts.Villarreal, J. C., & Renner, S. S. (2012) Hornwort pyrenoids, carbon-concentrating structures, evolved and were lost at least five times during the last 100 million years. ''Proceedings of the National Academy of Sciences'',109(46), 1873-1887. Pyrenoids are associated with the operation of a carbon-concentrating mechanism (CCM). Their main function is to act as centres of carbon dioxide (CO2) fixation, by generating and maintaining a CO2 rich environment around the photosynthetic enzyme ribulose-1,5-bisphosphate carboxylase/oxygenase (RuBisCO). Pyrenoids therefore seem to have a role analogous to that of carboxysomes in cyanobacteria. Algae are restricted to aqueous env ...
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Phymatoceros
''Phymatoceros'' is the only genus in the hornwort family Phymatocerotaceae and order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ... Phymatocerotales. It includes only two species. References External links Hornworts Bryophyte genera {{Bryophyte-stub ...
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Chloroplast
A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, and stores it in the energy-storage molecules ATP and NADPH while freeing oxygen from water in the cells. The ATP and NADPH is then used to make organic molecules from carbon dioxide in a process known as the Calvin cycle. Chloroplasts carry out a number of other functions, including fatty acid synthesis, amino acid synthesis, and the immune response in plants. The number of chloroplasts per cell varies from one, in unicellular algae, up to 100 in plants like ''Arabidopsis'' and wheat. A chloroplast is characterized by its two membranes and a high concentration of chlorophyll. Other plastid types, such as the leucoplast and the chromoplast, contain little chlorophyll and do not carry out photosynthesis. Chloroplasts are highly dynamic—they circulat ...
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Thallus
Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms were previously known as the thallophytes, a polyphyletic group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thallodal, thalliform, thalline, or thallose. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the tissues into organs. Even though thalli do not have organized and distinct parts (leaves, roots, and stems) as do the vascular plants, they may have analogous structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has vascular tissue. In exceptional cases such as the Lemnoideae, where ...
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