Heteronectes
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Heteronectes
''Heteronectes chaneti'' is a fossil fish which has been identified as a primitive flatfish, dating to the early Eocene (Lutetian stage) of France. ''Heteronectes'' is reported to be a transitional fossil. In a typical modern flatfish, the head is asymmetric with both eyes on one side of the head. In ''Heteronectes'', the transition from the typical symmetric head of a vertebrate is incomplete, with one eye positioned near the top of the head, very similar (but less so) to its Italian relative '' Amphistium''. The rest of the skeleton also has some primitive features in common with other Percomorph groups, but absent in living flatfishes.Friedman M (2008). “The evolutionary origin of flatfish asymmetry”, ''Nature'' 454(7201): p. 209-212: The condition in modern, bottom-dwelling flatfish with both eyes on the same side of the head was cited by St. George Jackson Mivart as difficult to imagine how it could have evolved in a gradual fashion by natural selection, as proposed ...
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Transitional Fossil
A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation. Because of the incompleteness of the fossil record, there is usually no way to know exactly how close a transitional fossil is to the point of divergence. Therefore, it cannot be assumed that transitional fossils are direct ancestors of more recent groups, though they are frequently used as models for such ancestors. In 1859, when Charles Darwin's ''On the Origin of Species'' was first published, the fossil record was poorly known. Darwin described the perceived lack of transitional fossils as "the most obvious and gravest objection which can be ...
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Amphistium
''Amphistium paradoxum'' (from el, ἀμφί , 'on both sides', el, ιστίον 'sail', and el, παράδοξος 'extraordinary'), the only species classified under the genus ''Amphistium'', is a fossil fish which has been identified as a Paleogene relative of the flatfish, and as a transitional fossil. In a typical modern flatfish, the head is asymmetric with both eyes on one side of the head. In ''Amphistium'', the transition from the typical symmetric head of a vertebrate is incomplete, with one eye placed near the top of the head. ''Amphistium'' is among the many fossil fish species known from the Monte Bolca ''Lagerstätte'' of Lutetian Italy. ''Heteronectes ''Heteronectes chaneti'' is a fossil fish which has been identified as a primitive flatfish, dating to the early Eocene (Lutetian stage) of France. ''Heteronectes'' is reported to be a transitional fossil. In a typical modern flatfish, the head ...'' is a related, and very similar fossil from a slightly e ...
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Transitional Fossils
A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation. Because of the incompleteness of the fossil record, there is usually no way to know exactly how close a transitional fossil is to the point of divergence. Therefore, it cannot be assumed that transitional fossils are direct ancestors of more recent groups, though they are frequently used as models for such ancestors. In 1859, when Charles Darwin's ''On the Origin of Species'' was first published, the fossil record was poorly known. Darwin described the perceived lack of transitional fossils as "the most obvious and gravest objection which can be ...
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Flatfish
A flatfish is a member of the Ray-finned fish, ray-finned demersal fish order (biology), order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migrating through or around the head during development. Some species face their left sides upward, some face their right sides upward, and others face either side upward. Many important food fish are in this order, including the flounders, sole (fish), soles, turbot, plaice, and halibut. Some flatfish can camouflage themselves on the ocean floor. Taxonomy Over 800 described species are placed into 16 families. Broadly, the flatfishes are divided into two suborders, Psettodoidei and Pleuronectoidei, with > 99% of the species diversity found within the Pleuronectoidei. The largest families are Soleidae, Bothidae and Tonguefish, Cynoglossidae with more than 150 species each. There also exist two monotypic families (P ...
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Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope Carbon-13, 13C in the atmosphere was exceptionally low in comparison with the more common isotope Carbon-12, 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Popigai impact structure, Siberia and in what is now ...
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Lutetian
The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Middle Eocene Subepoch. Stratigraphic definition The Lutetian was named after Lutetia, the Latin name for the city of Paris. The Lutetian Stage was introduced in scientific literature by French geologist Albert de Lapparent in 1883 and revised by A. Blondeau in 1981. The base of the Lutetian Stage is at the first appearance of the nanofossil ''Blackites inflatus'', according to an official reference profile (GSSP) established in 2011. Of two candidates located in Spain, the Gorrondatxe section was chosen.See thwebsite of Eustoquio Molinafor these candidates. The top of the Lutetian (the base of the Bartonian) is at the first appearance of calcareous nanoplankton species ''Reticulofenestra reticulata''. The Lutetian overlaps with the Geisel ...
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Percomorpha
Percomorpha () is a large clade of ray-finned fish that includes the tuna, seahorses, gobies, cichlids, flatfish, wrasse, perches, anglerfish, and pufferfish. Evolution Percomorpha are the most diverse group of teleost fish today. Teleosts, and percomorphs in particular, thrived during the Cenozoic era. Fossil evidence shows that there was a major increase in size and abundance of teleosts immediately after the mass extinction event at the Cretaceous-Paleogene boundary ca. 65 Ma ago. Phylogeny External relationships The two cladograms below are based on Betancur-R ''et al.'', 2017. Percomorphs are a clade of teleost fishes. The first cladogram shows the interrelationships of percomorphs with other living groups of teleosts. Internal relationships The following cladogram shows the evolutionary relationships of the various groups of extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, ...
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Natural Selection
Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Charles Darwin popularised the term "natural selection", contrasting it with selective breeding, artificial selection, which in his view is intentional, whereas natural selection is not. Genetic diversity, Variation exists within all populations of organisms. This occurs partly because random mutations arise in the genome of an individual organism, and their offspring can inherit such mutations. Throughout the lives of the individuals, their genomes interact with their environments to cause variations in traits. The environment of a genome includes the molecular biology in the Cell (biology), cell, other cells, other individuals, populations, species, as well as the abiotic environment. Because individuals with certain variants of the trait tend ...
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Charles Darwin
Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended from a common ancestor is now generally accepted and considered a fundamental concept in science. In a joint publication with Alfred Russel Wallace, he introduced his scientific theory that this branching pattern of evolution resulted from a process he called natural selection, in which the struggle for existence has a similar effect to the artificial selection involved in selective breeding. Darwin has been described as one of the most influential figures in human history and was honoured by burial in Westminster Abbey. Darwin's early interest in nature led him to neglect his medical education at the University of Edinburgh; instead, he helped to investigate marine invertebrates. His studies at the University of Cambridge's Christ's Col ...
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Saltation (biology)
In biology, saltation () is a sudden and large mutational change from one generation to the next, potentially causing single-step speciation. This was historically offered as an alternative to Darwinism. Some forms of mutationism were effectively saltationist, implying large discontinuous jumps. Speciation, such as by polyploidy in plants, can sometimes be achieved in a single and in evolutionary terms sudden step. Evidence exists for various forms of saltation in a variety of organisms. History Prior to Charles Darwin most evolutionary scientists had been saltationists. Jean-Baptiste Lamarck was a gradualist but similar to other scientists of the period had written that saltational evolution was possible. Étienne Geoffroy Saint-Hilaire endorsed a theory of saltational evolution that "monstrosities could become the founding fathers (or mothers) of new species by instantaneous transition from one form to the next." Geoffroy wrote that environmental pressures could produce sudden ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the ...
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Pleuronectiformes
A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migrating through or around the head during development. Some species face their left sides upward, some face their right sides upward, and others face either side upward. Many important food fish are in this order, including the flounders, soles, turbot, plaice, and halibut. Some flatfish can camouflage themselves on the ocean floor. Taxonomy Over 800 described species are placed into 16 families. Broadly, the flatfishes are divided into two suborders, Psettodoidei and Pleuronectoidei, with > 99% of the species diversity found within the Pleuronectoidei. The largest families are Soleidae, Bothidae and Cynoglossidae with more than 150 species each. There also exist two monotypic families (Paralichthodidae and Oncopteridae). Some families are ...
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