Hesperotestudo Alleni
''Hesperotestudo'' ("Western turtle") is an extinct genus of tortoise native to North and Central America from the Oligocene to the Late Pleistocene. Species of ''Hesperotesudo'' varied widely in size, with the largest unnamed species from El Salvador reaching in carapace length, larger than that of extant giant tortoises. Historically considered a subgenus of ''Geochelone'', it is now considered to be distantly related to that genus. Its closest relatives are the extant ''Gopherus'' (gopher tortoises) and the extinct '' Stylemys.'' The ancestor of the three genera arrived in North America during the Early Eocene. The bodies of ''Hesperotesudo'' species were extensively covered with large dermal ossicles, which in life were covered in keratin. It has been suggested that species of ''Hesperotestudo'' were relatively tolerant of cold weather. ''Hesperotestudo'' became extinct at the end of the Pleistocene and beginning of the Holocene co-incident with the arrival of the first huma ...
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Extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. More than 99% of all species that ever lived on Earth, amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryote globally, and possibly many times more if microorganisms, like bacteria, are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, dodos, m ...
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Before Present
Before Present (BP) years, or "years before present", is a time scale used mainly in archaeology, geology and other scientific disciplines to specify when events occurred relative to the origin of practical radiocarbon dating in the 1950s. Because the "present" time changes, standard practice is to use 1 January 1950 as the commencement date (epoch) of the age scale. The abbreviation "BP" has been interpreted retrospectively as "Before Physics", which refers to the time before nuclear weapons testing artificially altered the proportion of the carbon isotopes in the atmosphere, which scientists must now account for. In a convention that is not always observed, many sources restrict the use of BP dates to those produced with radiocarbon dating; the alternative notation RCYBP stands for the explicit "radio carbon years before present". Usage The BP scale is sometimes used for dates established by means other than radiocarbon dating, such as stratigraphy. This usage differs from t ...
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Pliocene Turtles
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58See the 2014 version of the ICS geologic time scale
million years ago. It is the second and most recent epoch of the Period in the . The Pliocene follows the Epoch and is followed by the

Miocene Turtles
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Hesperotestudo
''Hesperotestudo'' ("Western turtle") is an extinct genus of tortoise native to North and Central America from the Oligocene to the Late Pleistocene. Species of ''Hesperotesudo'' varied widely in size, with the largest unnamed species from El Salvador reaching in carapace length, larger than that of extant giant tortoises. Historically considered a subgenus of ''Geochelone'', it is now considered to be distantly related to that genus. Its closest relatives are the extant ''Gopherus'' (gopher tortoises) and the extinct '' Stylemys.'' The ancestor of the three genera arrived in North America during the Early Eocene. The bodies of ''Hesperotesudo'' species were extensively covered with large dermal ossicles, which in life were covered in keratin. It has been suggested that species of ''Hesperotestudo'' were relatively tolerant of cold weather. ''Hesperotestudo'' became extinct at the end of the Pleistocene and beginning of the Holocene co-incident with the arrival of the first huma ...
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Burdigalian
The Burdigalian is, in the geologic timescale, an age (geology), age or stage (stratigraphy), stage in the early Miocene. It spans the time between 20.43 ± 0.05 annum, Ma and 15.97 ± 0.05 Ma (million years ago). Preceded by the Aquitanian (stage), Aquitanian, the Burdigalian was the first and longest warming period of the MioceneEdward Petuch, Ph.D. Florida Atlantic University, Department of Geosciences. and is succeeded by the Langhian. Stratigraphic definition The name Burdigalian comes from ''Burdigala'', the Latin name for the city of Bordeaux, France. The Burdigalian Stage was introduced in scientific literature by Charles Depéret in 1892. The base of the Burdigalian is at the first appearance of foram species ''Globigerinoides altiaperturus'' and the top of magnetic chronozone C6An. , an official GSSP for the Burdigalian had not yet been assigned. The top of the Burdigalian (the base of the Langhian) is defined by the first appearance of foram species ''Praeorbulina gl ...
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Oakville Formation
The Oakville Formation is a geologic formation in Texas. It preserves fossils dating back to the Neogene period. See also * List of fossiliferous stratigraphic units in Texas * Paleontology in Texas Paleontology in Texas refers to paleontological research occurring within or conducted by people from the U.S. state of Texas. Author Marian Murray has remarked that "Texas is as big for fossils as it is for everything else." Some of the most imp ... References * Neogene geology of Texas {{Neogene-stub ...
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Zanclean
The Zanclean is the lowest stage or earliest age on the geologic time scale of the Pliocene. It spans the time between 5.332 ± 0.005 Ma (million years ago) and 3.6 ± 0.005 Ma. It is preceded by the Messinian Age of the Miocene Epoch, and followed by the Piacenzian Age. The Zanclean can be correlated with regionally used stages, such as the Opoitian of New Zealand, and the Tabianian or Dacian of Central Europe. It also corresponds to the late Hemphillian to mid-Blancan North American Land Mammal Ages. In California, the Zanclean roughly corresponds to the middle part of the Delmontian stage. Definition The Zanclean Stage was introduced by Giuseppe Seguenza in 1868. It is named after ''Zancle'', the pre-Roman name for the Italian city of Messina on Sicily. The base of the Zanclean (and the Pliocene Series) lies with the top of magnetic chronozone Cr3 (about 100,000 years before the Thvera normal subchronozone C3n.4n). The base is also close to the extinction level of the calc ...
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Tortonian
The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million years ago). It follows the Serravallian and is followed by the Messinian. The Tortonian roughly overlaps with the regional Pannonian Stage of the Paratethys timescale of Central Europe. It also overlaps the upper Astaracian, Vallesian and lower Turolian European land mammal ages, the upper Clarendonian and lower Hemphillian North American land mammal ages and the upper Chasicoan and lower Huayquerian South American land mammal ages. Definition The Tortonian was introduced by Swiss stratigrapher Karl Mayer-Eymar in 1858. It was named after the Italian city of Tortona in the region Piedmont. The base of the Tortonian Stage is at the last common appearance of calcareous nanoplankton '' Discoaster kugleri'' and planktonic foram '' Globigerinoides subquadratus''. It is also associated with the short normal polarized magneti ...
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Hemphillian
The Hemphillian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 10,300,000 to 4,900,000 years BP. It is usually considered to overlap the Tortonian age of the Late Miocene and Zanclean age of the Early Pliocene. The Hemphillian is preceded by the Clarendonian and followed by the Blancan The Blancan North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 4,750,000 to 1,806,000 years Before Present, BP, a period of .

Ogallala Formation
The Ogallala Formation is a Miocene to early Pliocene geologic formation in the central High Plains of the western United States and the location of the Ogallala Aquifer. In Nebraska and South Dakota it is also classified as the Ogallala Group. Notably, it records the North American Land Mammal Ages (NALMAs) Hemphillian, Clarendonian, and Barstovian. It also includes an excellent record of grass seeds and other plant seeds, which can be used for biostratigraphic dating within the formation. The Ogallala Formation outcrops of Lake Meredith National Recreation Area preserve fish fossils. Similar specimens from the same unit are found at Alibates Flint Quarries National Monument in Texas. Development At the beginning of the Rocky Mountains uplift, the final floor of the Western Interior Seaway was elevated several thousand feet (thousands of meters) above sea level. The previous pattern of erosion of material from the northeast into the midcontinent was largely reversed so that m ...
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