Hebediscidae
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Hebediscidae
The Hebediscidae Kobayashi, 1944, are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Atdabanian to Toyonian). They are small or very small, and have a thorax of two or three segments. The Hebediscidae include five genera (see box). Taxonomy The Hebediscidae probably descended from the Tsunydiscidae and gave rise to the Weymouthiidae. cited in Description Like all Agnostida, the Hebediscidae are diminutive and the headshield (or cephalon) and tailshield (or pygidium) are of approximately the same size (or isopygous) and outline. In the Hebediscidae, the central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) is wide at its rear end, has parallel sides or tapers forward, and without t ...
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Parapagetia
''Parapagetia'' is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the Botomian stage, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the Cambrian Period. Distribution * ''P. limbata'' is known from the Lower Cambrian of the Russian Federation (Botomian, Sanashtykgol, Gorno-Altayskaya, Katun River). * ''P. plana'' has been found in the Lower Cambrian of the Russian Federation (Botomian, Sanashtykgol, Gorno-Altayskaya, Ul’men’River). Description Like all Agnostida, the Hebediscidae are diminutive and the headshield (or cephalon) and tailshield (or pygidium The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is compo ...) are of approximately the same size (or isopygous) and outline. ...
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Eodiscina
Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period (late Atdabanian) and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea. Taxonomy The Eodiscina are mostly considered the more primitive suborder of the Agnostida, and the Agnostina the more advanced. Some scholars do not consider the Agnostina true trilobites, and consequently rejected the idea that they were related to the Eodiscina. Consequently, these scientists have proposed to elevate the group to ordinal level, which would thus be called Eodiscida Kobayashi, 1939. Origin The oldest known eodiscoid is ''Tsunyidiscus''. The glabella of ''Tsunyidiscus'' is extremely similar to that of ''Dipharus clarki'', and distinct from all other eodiscoids. ''D. clarki'' is thought to represent an immature stage of the redlichioid ''Bu ...
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Tsunydiscidae
''Tsunyidiscus'' is a trilobite belonging to the Suborder Eodiscina. ''Tsunyidiscus'' appeared near the end of the Lower Cambrian, during the late Atdabanian stage of geologic time and some collections suggest it may have survived into the Botomian. The genus is very small (up to 7mm), oculate and isopypous with a narrow dome-shaped glabella and a narrow bullet-shaped pygidial axis. Thorax consists of three segments. ''Tsunyidiscus'' is the only genus currently attributed to the family Tsunyidiscidae. Description Like other Agnostida the exoskeleton of ''Tsunyidiscus'' is diminutive and isopygous with 3 fulcrate thoracic segments. The cephalon has a strongly parabolic outline and maximum width (tr.) usually anterior to the genal angles. Glabella extremely narrow, lateral glabellar furrows usually obscure, with a rounded and expanded frontal glabellar lobe. The occipital ring (LO) is at least as long as L1, usually expanded laterally, and may bear a sharp posteriorly directed s ...
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Agnostida
Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician. Systematics The Agnostida are divided into two suborders — Agnostina and Eodiscina — which are then subdivided into a number of families. As a group, agnostids are isopygous, meaning their pygidium is similar in size and shape to their cephalon. Most agnostid species were eyeless. The systematic position of the order Agnostida within the class Trilobita remains uncertain, and there has been continuing deb ...
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Delgadella
''Delgadella'' is a diminutive trilobite that lived during the late Lower Cambrian (Atdabanian to Botomian, over 500 million years ago) and has been found in Russia (Siberian Platform, Altay Mountains), Mongolia, Spain, Italy (Sardinia), Portugal, Morocco and Canada (Newfoundland). It can be recognized by its strongly effaced headshield and tailshield, with narrow but distinct furrows and borders along its margins, and three thorax segments. Description The headshield (or cephalon) is convex, and axial furrow that surrounds the central area (or glabella) almost obsolete, particularly on the external surface. The glabella has no transverse furrows. The border furrow is distinct and wide anteriorly, and the border distinct and narrow. The eye lobe (or palpebral lobe) is poorly defined. The free cheeks (or librigenae) are long. The thorax has three segments. The tailshield (or pygidium The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some othe ...
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Dicerodiscus
''Dicerodiscus'' is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the early part of the Botomian stage, in China (Guizhou and Sichuan). Four species have been assigned to it. ''Dicerodiscus'' is unique for an eodiscoid in having conspicuous and curved spines that are attached anteriorly, and at their base are directed outward perpendicular to the midline, before gradually bending further backwards. Distribution * ''D. tsunyiensis'' is known from the Lower Cambrian of China (Canglangpuan, Guizhou). * ''D. pertenus'' has been found in the Lower Cambrian (Qiongzhusian, Sichuan). Description Like all Agnostida, ''Dicerodiscus'' is diminutive and the headshield (or cephalon) and tailshield (or pygidium The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, th ...
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Weymouthiidae
The Weymouthiidae Kobayashi 1943 are an extinct family of eodiscinid agnostid trilobites. They lived during the late Lower Cambrian and earliest Middle Cambrian (Botomian to Delamaran) in the so-called ''Olenellus''- and ''Eokochaspis''-zones in the former paleocontinents of Laurentia, Avalonia, Gondwana. The Weymouthiidae are all blind and lack free cheeks. Taxonomy Ancestors The Weymouthiidae have developed from a stock within the Hebediscidae The Hebediscidae Kobayashi, 1944, are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Atdabanian to Toyonian). They are small or very small, and have a thorax of two or three segments. The Hebediscida .... Descendants The Weymouthiidae are a paraphyletic family because the Agnostina suborder is nested within it, particularly in the clade that further consists of the genera ''Mallagnostus'', ''Chelediscus'', ''Tannudiscus'' and ''Jinghediscus''. The trend in the Weymouthiid ...
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Tchernyshevioides
''Tchernyshevioides'' is a diminutive trilobite that lived during the early Middle Cambrian ( Amgaian) and has been found in the Russian Federation (''Pseudanomocarina''-zone, Sulyukta region, Tian-Shan) and the Jbel Wawrmast Formation of Morocco.Whittington, H. B. et al. Part O, Treatise on Invertebrate Paleontology. Revised, Volume 1 – Trilobita – Introduction, Order Agnostida, Order Redlichiida. 1997 Etymology ''Tchernyshevioides'' has been named in honor of the Russian paleontologist N.E. Tchernysheva. Description The central raised area (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) of the headshield (or cephalon) is long and reaches the anterior border furrow. The occipital ring is well defined and carries a long posterodorsal spine. Otherwise th ...
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Cambrian Trilobites
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian bi ...
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Trilobite Families
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The stud ...
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Pygidium
The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is composed of fused body segments, sometimes with a tail, and separated from thoracic segments by an articulation.Shultz, J.W. (1990). Evolutionary Morphology And Phylogeny of Arachnida. Cladistics 6: 1–38. Chelicerates In arachnids, the pygidium is formed by reduction of the last three opisthosomal segments to rings where there is no distinction between tergites and sternites. A pygidium is present in Palpigradi, Amblypygi, Thelyphonida, Schizomida, Ricinulei and in the extinct order Trigonotarbida. It is also present in early fossil representatives of horseshoe crabs. Trilobites In trilobites, the pygidium can range from extremely small (much smaller than the head, or cephalon) to larger than the cephalon. They can be smooth, as in order ...
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Pygidium
The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is composed of fused body segments, sometimes with a tail, and separated from thoracic segments by an articulation.Shultz, J.W. (1990). Evolutionary Morphology And Phylogeny of Arachnida. Cladistics 6: 1–38. Chelicerates In arachnids, the pygidium is formed by reduction of the last three opisthosomal segments to rings where there is no distinction between tergites and sternites. A pygidium is present in Palpigradi, Amblypygi, Thelyphonida, Schizomida, Ricinulei and in the extinct order Trigonotarbida. It is also present in early fossil representatives of horseshoe crabs. Trilobites In trilobites, the pygidium can range from extremely small (much smaller than the head, or cephalon) to larger than the cephalon. They can be smooth, as in order ...
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