Hapalodectes
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Hapalodectes
''Hapalodectes'' (literal translation 'soft biter'; from grc, ἁπαλός ('soft, tender') and ('biter')) is an extinct genus of otter-like mesonychids from the Late Paleocene to Early Eocene, some 55million years ago. Although the first fossils were found in the Eocene strata of Wyoming, the genus originated in Mongolia, as the oldest species is ''H. dux'', which was found in Late Paleocene strata in the Naran Bulak Formation. The genus was once suggested to be related to the Archaeoceti, such as '' Pakicetus'', due to numerous similarities between the skull and tooth anatomies of the two genera. Now, however, ''Hapalodectes'' and other mesonychians are thought to be related to basal artiodactyls, while the Archaeoceti are now determined to be descended from more derived artiodactyls, like '' Indohyus'', which are related to hippopotamuses and anthracothere Anthracotheriidae is a paraphyletic family of extinct, hippopotamus-like artiodactyl ungulates related t ...
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Pachyaena
''Pachyaena'' (literally, "thick hyena") was a genus of heavily built, relatively short-legged mesonychids, early Cenozoic mammals that evolved before the origin of either modern hoofed animals or carnivores, and combined characteristics similar to both. The genus likely originated from Asia and spread to Europe, and from there to North America across a land bridge in what is now the North Atlantic ocean. Various described species of ''Pachyaena'' ranged in size from a coyote to a bear. However, recent work indicates that ''Pachyaena'' is paraphyletic, with ''P. ossifraga'' being closer to '' Synoplotherium'', '' Harpagolestes'' and ''Mesonyx'' than to ''P. gigantea''. Unlike many mesonychids, ''Pachyaena'' is known from skeletal material in addition to skulls and jaws. Analysis of three wolf- to bear-sized species from the early Eocene of Wyoming (Willwood formation) indicates they all had many adaptations for running, including paraxonic compressed feet with a vestigial first ...
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Late Paleocene
The Thanetian is, in the International Commission on Stratigraphy, ICS Geologic timescale, the latest age (geology), age or uppermost stage (stratigraphy), stratigraphic stage of the Paleocene epoch (geology), Epoch or series (stratigraphy), Series. It spans the time between . The Thanetian is preceded by the Selandian Age and followed by the Ypresian Age (part of the Eocene). The Thanetian is sometimes referred to as the Late Paleocene. Stratigraphic definition The Thanetian was established by Switzerland, Swiss geologist Eugène Renevier in 1873. The Thanetian is named after the Thanet Formation, the oldest Cenozoic deposit of the London Basin, which was first identified in the area of Kent (southern England) known as the Isle of Thanet. The base of the Thanetian Stage is laid at the base of magnetic chronozone C26n. The references profile (Global Boundary Stratotype Section and Point) is in the Zumaia section (43° 18'N, 2° 16'W) at the beach of Itzurun, Pais Vasco, northern ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Ypresian Extinctions
In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian is consistent with the lower Eocene. Events The Ypresian Age begins during the throes of the Paleocene–Eocene Thermal Maximum (PETM). The Fur Formation in Denmark, the Messel shales in Germany, the Oise amber of France and Cambay amber of India are of this age. The Eocene Okanagan Highlands are an uplands subtropical to temperate series of lakes from the Ypresian. Stratigraphic definition The Ypresian Stage was introduced in scientific literature by Belgian geologist André Hubert Dumont in 1850. The Ypresian is named after the Flemish city of Ypres in Belgium (spelled ''Ieper'' in Dutch). The definitions of the original stage were totally different from the modern ones. The Ypresian shares its name with the Belgian Ieper Group (French: ...
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Eocene Mammals
The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope 13C in the atmosphere was exceptionally low in comparison with the more common isotope 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Siberia and in what is now Chesapeake Bay. As with other geologic periods, the strata that define the start and end of the ...
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Paleocene Mammals
The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''palaiós'' meaning "old" and the Eocene Epoch (which succeeds the Paleocene), translating to "the old part of the Eocene". The epoch is bracketed by two major events in Earth's history. The K–Pg extinction event, brought on by an asteroid impact and possibly volcanism, marked the beginning of the Paleocene and killed off 75% of living species, most famously the non-avian dinosaurs. The end of the epoch was marked by the Paleocene–Eocene Thermal Maximum (PETM), which was a major climatic event wherein about 2,500–4,500 gigatons of carbon were released into the atmosphere and ocean systems, causing a spike in global temperatures and ocean acidification. In the Paleocene, the continents of the Northern Hemisphere were still connected via som ...
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Mesonychids
Mesonychia ("middle claws") is an extinct taxon of small- to large-sized carnivorous ungulates related to artiodactyls. Mesonychids first appeared in the early Paleocene, went into a sharp decline at the end of the Eocene, and died out entirely when the last genus, ''Mongolestes'', became extinct in the early Oligocene. In Asia, the record of their history suggests they grew gradually larger and more predatory over time, then shifted to scavenging and bone-crushing lifestyles before the group became extinct. Mesonychids probably originated in China, where the most primitive mesonychid, ''Yangtanglestes'', is known from the early Paleocene. They were also most diverse in Asia, where they occur in all major Paleocene faunas. Since other predators, such as creodonts and Carnivora, were either rare or absent in these animal communities, mesonychids most likely dominated the large predator niche in the Paleocene of eastern Asia. One genus, ''Dissacus'', had successfully spread to Eur ...
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Anthracothere
Anthracotheriidae is a paraphyletic family of extinct, hippopotamus-like artiodactyl ungulates related to hippopotamuses and whales. The oldest genus, '' Elomeryx'', first appeared during the middle Eocene in Asia. They thrived in Africa and Eurasia, with a few species ultimately entering North America during the Oligocene. They died out in Europe and Africa during the Miocene, possibly due to a combination of climatic changes and competition with other artiodactyls, including pigs and true hippopotamuses. The youngest genus, '' Merycopotamus'', died out in Asia during the late Pliocene. The family is named after the first genus discovered, '' Anthracotherium'', which means "coal beast", as the first fossils of it were found in Paleogene-aged coal beds in France. Fossil remains of the anthracothere genus were discovered by the Harvard University and Geological Survey of Pakistan joint research project (Y-GSP) in the well-dated middle and late Miocene deposits of the Pothohar Pla ...
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Hippopotamus
The hippopotamus ( ; : hippopotamuses or hippopotami; ''Hippopotamus amphibius''), also called the hippo, common hippopotamus, or river hippopotamus, is a large semiaquatic mammal native to sub-Saharan Africa. It is one of only two extant species in the family Hippopotamidae, the other being the pygmy hippopotamus (''Choeropsis liberiensis'' or ''Hexaprotodon liberiensis''). Its name comes from the ancient Greek for "river horse" (). Aside from elephants and rhinos, the hippopotamus is the largest land mammal. It is also the largest extant land artiodactyl. Despite their physical resemblance to pigs and other terrestrial even-toed ungulates, the closest living relatives of the hippopotamids are cetaceans (whales, dolphins, porpoises, etc.), from which they diverged about 55 million years ago. Hippos are recognisable for their barrel-shaped torsos, wide-opening mouths with large canine tusks, nearly hairless bodies, pillar-like legs, and large size: adults average ...
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Indohyus
''Indohyus'' is an extinct genus of digitigrade artiodactyls known from Eocene fossils in Asia. This small chevrotain-like animal found in the Himalayas is one of the earliest known non-cetacean ancestors of whales. Discovery The fossils were discovered among rocks that had been collected more than 30 years ago in Kashmir by the Indian geologist A. Ranga Rao who found a few teeth and parts of a jawbone, but when he died many rocks had yet to be broken open. Ranga Rao's widow gave the rocks to Hans Thewissen, who was working on them when his technician accidentally broke one of the skulls they had found and Thewissen recognised the ear structure of the auditory bulla, formed from the ectotympanic bone in a shape which is highly distinctive, found only in the skulls of cetaceans both living and extinct, including ''Pakicetus''. Paleobiology About the size of a raccoon or domestic cat, this omnivorous pig-like creature shared some of the traits of whales, and showed signs of adap ...
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Derived (phylogenetics)
In phylogenetics, a primitive (or ancestral) character, trait, or feature of a lineage or taxon is one that is inherited from the common ancestor of a clade (or clade group) and has undergone little change since. Conversely, a trait that appears ''within'' the clade group (that is, is present in any subgroup within the clade but not all) is called advanced or derived. A clade is a group of organisms that consists of a common ancestor and all its lineal descendants. A primitive trait is the original condition of that trait in the common ancestor; advanced indicates a notable change from the original condition. These terms in biology contain no judgement about the sophistication, superiority, value or adaptiveness of the named trait. "Primitive" in biology means only that the character appeared first in the common ancestor of a clade group and has been passed on largely intact to more recent members of the clade. "Advanced" means the character has evolved within a later subgroup ...
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Artiodactyl
The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, odd-toed ungulates bear weight on an odd number of the five toes. Another difference between the two is that many other even-toed ungulates (with the exception of Suina) digest plant cellulose in one or more stomach chambers rather than in their intestine as the odd-toed ungulates do. Cetaceans (whales, dolphins, and porpoises) evolved from even-toed ungulates, and are therefore often classified under the same taxonomic branch because a species cannot outgrow its evolutionary ancestry; some modern taxonomists combine the two under the name Cetartiodactyla , while others opt to include cetaceans in the already-existing Artiodactyla. The roughly 270 land-based even-toed ungulate species include pigs, peccaries, hippopota ...
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