Halisaurinae
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Halisaurinae
The Halisaurinae are a subfamily of mosasaurs, a group of Late Cretaceous marine lizards. They were small to medium-sized, ranging from just under 3 meters in ''Eonatator sternbergi'' to as much as 8 or 9 meters in ''Pluridens serpentis''. They tended to have relatively slender jaws and small, numerous teeth, suggesting a diet of small fish and other prey. Although the skeleton is primitive compared to other Mosasauridae in many respects, halisaurines had the distinctive hypocercal tail of other mosasaurids suggesting good swimming ability, and they persisted alongside other mosasaurs until the end of the Cretaceous. The earliest known remains of halisaurines occur in rocks of Santonian age and the subfamily persists until the latest Maastrichtian. Halisaurines are known from North and South America, Europe, Asia and Africa, indicating a more or less global distribution in the Late Cretaceous. Four genera are currently recognized: ''Eonatator'', ''Halisaurus'', '' Phosphorosaur ...
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Halisaurus Arambourgi
''Halisaurus'' is an extinct genus of marine reptile belonging to the mosasaur family. The holotype, consisting of an angular and a basicranium fragment discovered near Hornerstown, New Jersey, already revealed a relatively unique combination of features and prompted a new genus to be described. It was named by Othniel Charles Marsh in 1869 and means "ocean lizard". It was renamed by Marsh to ''Baptosaurus'' in 1870, since he believed the name to already be preoccupied by the fish ''Halosaurus''. According to modern rules, a difference of a letter is enough and the substitute name is unneeded, making "''Baptosaurus''" a junior synonym. Since its description, more complete remains have been uncovered from fossil deposits throughout the world with particularly complete remains found in Morocco and the United States. The genus remains a key taxon in mosasaur systematics due to its unique set of features and as the most complete representative of its subfamily, the Halisaurinae. W ...
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Halisaurus
''Halisaurus'' is an extinct genus of marine reptile belonging to the mosasaur family. The holotype, consisting of an angular and a basicranium fragment discovered near Hornerstown, New Jersey, already revealed a relatively unique combination of features and prompted a new genus to be described. It was named by Othniel Charles Marsh in 1869 and means "ocean lizard". It was renamed by Marsh to ''Baptosaurus'' in 1870, since he believed the name to already be preoccupied by the fish ''Halosaurus''. According to modern rules, a difference of a letter is enough and the substitute name is unneeded, making "''Baptosaurus''" a junior synonym. Since its description, more complete remains have been uncovered from fossil deposits throughout the world with particularly complete remains found in Morocco and the United States. The genus remains a key taxon in mosasaur systematics due to its unique set of features and as the most complete representative of its subfamily, the Halisaurinae. ...
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Eonatator
''Eonatator'' is an extinct genus of marine lizard belonging to the mosasaur family. It is a close relative of ''Halisaurus'', and part of the same subfamily, the Halisaurinae. It is known from the Late Cretaceous of North America, Colombia and Sweden. Originally, this taxon was included within ''Halisaurus'', but was placed in its own genus, which also led to the subfamily Halisaurinae being created for the two genera. Discovery and naming ''Eonatator'' is known from the Smoky Hill Chalk Member of the Niobrara Chalk Formation ( Late Coniacian to Early Campanian) of Kansas, from the Eutaw Formation (Santonian) and Mooreville Chalk Formation (Selma Group; Santonian-Lower Campanian) of Alabama (United States), from the Kristianstad Basin of southern Sweden (late early Campanian), and the unit Nivel de Lutitas y Arenas (Campanian) of the Olini Group in La Mesa, Colombia. The name ''Eonatator'' means "dawn swimmer" (Greek ''eos'' = dawn + Latin ''natator'' = swimmer). Originally ...
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Phosphorosaurus
''Phosphorosaurus'' ("phosphate lizard") is an extinct genus of marine lizard belonging to the mosasaur family. ''Phosphorosaurus'' is classified within the Halisaurinae subfamily alongside the genera '' Pluridens'', ''Eonatator'' and ''Halisaurus''. Stratigraphically, ''Phosphorosaurus'' only occurs in the Maastrichtian stage of the Late Cretaceous. Although treated as synonymous with ''Halisaurus'' in the past, recent studies recognize it as valid. Two species are known, ''Phosphorosaurus ortliebi'' from the Craie de Ciply Formation in Belgium, and ''P. ponpetelegans'' from the Hakobuchi Formation of Hokkaido in Japan. ''P. ponpetelegans'' is only known from the very earliest Maastrichtian, whilst ''P. ortliebi'' occurs throughout the Maastrichtian. Description With a length of around 3 m (10 ft), ''Phosphorosaurus'' was small compared to most other mosasaurs, but rather standard in size for a halisaurine. Analysis of ''Phosphorosaurus'' biology suggests that th ...
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Pluridens
''Pluridens'' ("many teeth") is an extinct genus of marine lizard belonging to the Mosasauridae. ''Pluridens'' is placed in the subfamily Halisaurinae with the genera ''Phosphorosaurus'', ''Eonatator'' and ''Halisaurus''.Konishi, Takuya; Caldwell, Michael W.; Nishimura, Tomohiro; Sakurai, Kazuhiko; Tanoue, Kyo (2015). "A new halisaurine mosasaur (Squamata: Halisaurinae) from Japan: the first record in the western Pacific realm and the first documented insights into binocular vision in mosasaurs". Journal of Systematic Palaeontology: 1–31. doi:10.1080/14772019.2015.1113447. Compared to related halisaurines, ''Pluridens'' had longer jaws with more teeth, and smaller eyes. It also grew large size, measuring long and perhaps over in some individuals. The jaws in some specimens are robust, and sometimes show injuries suggestive of combat. The jaws may have been used for fighting over mates or territories. ''Pluridens'' lived in the shallow seas of West Africa during the late Camp ...
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Mosasaur
Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes. Mosasaurs probably evolved from an extinct group of aquatic lizards known as aigialosaurs in the Earliest Late Cretaceous with 42 described genera. During the last 20 million years of the Cretaceous period (Turonian–Maastrichtian ages), with the extinction of the ichthyosaurs and pliosaurs, mosasaurs became the dominant marine predators. They themselves became extinct as a result of the K-Pg event at the end of the Cretaceous period, about 66 million years ago. Description Mosasaurs breathed air, were powerful swimmers, and were well-adapted to living in the warm, shallow inland seas prevalent during the Late Cretaceous period. Mosasaurs were so ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxillary bones are fused at the intermaxillary suture, forming the anterior nasal spine. This is similar to the mandible (lower jaw), which is also a fusion of two mandibular bones at the mandibular symphysis. The mandible is the movable part of the jaw. Structure In humans, the maxilla consists of: * The body of the maxilla * Four processes ** the zygomatic process ** the frontal process of maxilla ** the alveolar process ** the palatine process * three surfaces – anterior, posterior, medial * the Infraorbital foramen * the maxillary sinus * the incisive foramen Articulations Each maxilla articulates with nine bones: * two of the cranium: the frontal and ethmoid * seven of the face: the nasal, zygomatic, lacrimal, inferior n ...
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Sweden
Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, Finland to the east, and is connected to Denmark in the southwest by a bridgetunnel across the Öresund. At , Sweden is the largest Nordic country, the third-largest country in the European Union, and the fifth-largest country in Europe. The capital and largest city is Stockholm. Sweden has a total population of 10.5 million, and a low population density of , with around 87% of Swedes residing in urban areas in the central and southern half of the country. Sweden has a nature dominated by forests and a large amount of lakes, including some of the largest in Europe. Many long rivers run from the Scandes range through the landscape, primarily ...
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Moanasaurus
''Moanasaurus'' (From Māori ''moana'' "sea" and Greek ''sauros'' "lizard"; meaning "Sea Lizard") was a genus of mosasaur from the Late Cretaceous period. Its fossil remains have been discovered in the North Island of New Zealand. ''Moanasaurus'' was a very large mosasaurine known originally from a disarticulated skull, vertebrae, ribs and flipper bones. The skull measures in length, which shows that ''Moanasaurus'' was one of the largest in the subfamily of Mosasaurinae. Researchers argue that some Antarctic '' Mosasaurus'' remains (including a "large, fragmentaery skull") may be attributed to this genus. Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ... estimated its maximum adult size at in length and in body mass. See also * List of dinosaurs and other ...
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Premaxilla
The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has been usually termed as the incisive bone. Other terms used for this structure include premaxillary bone or ''os premaxillare'', intermaxillary bone or ''os intermaxillare'', and Goethe's bone. Human anatomy In human anatomy, the premaxilla is referred to as the incisive bone (') and is the part of the maxilla which bears the incisor teeth, and encompasses the anterior nasal spine and alar region. In the nasal cavity, the premaxillary element projects higher than the maxillary element behind. The palatal portion of the premaxilla is a bony plate with a generally transverse orientation. The incisive foramen is bound anteriorly and laterally by the premaxilla and posteriorly by the palatine process of the maxilla. It is formed from the ...
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Fibula
The fibula or calf bone is a leg bone on the lateral side of the tibia, to which it is connected above and below. It is the smaller of the two bones and, in proportion to its length, the most slender of all the long bones. Its upper extremity is small, placed toward the back of the head of the tibia, below the knee joint and excluded from the formation of this joint. Its lower extremity inclines a little forward, so as to be on a plane anterior to that of the upper end; it projects below the tibia and forms the lateral part of the ankle joint. Structure The bone has the following components: * Lateral malleolus * Interosseous membrane connecting the fibula to the tibia, forming a syndesmosis joint * The superior tibiofibular articulation is an arthrodial joint between the lateral condyle of the tibia and the head of the fibula. * The inferior tibiofibular articulation (tibiofibular syndesmosis) is formed by the rough, convex surface of the medial side of the lower end of the f ...
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