Globidentosuchus Brachyrostris
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Globidentosuchus Brachyrostris
''Globidentosuchus'' is an extinct genus of basal caimanine crocodylian known from the late Middle to Late Miocene of the Middle and the Upper Members of the Urumaco Formation at Urumaco, Venezuela. Its skull was very short and robust, with large units of spherical teeth used to break the shells of molluscs as part of its durophagus diet. It is thought to be one of the most basal Caimanines, even sharing some traits with alligatorids. Etymology The generic name ''Globidentosuchus'' is derived from the Latin roots ''globus'' meaning "sphere" and ''dens'' meaning "tooth", referring to the spherical teeth in the posterior skull, and Greek ''souchos'' meaning "crocodile" after its classification. The species name ''brachyrostris'' is derived from the Greek ''brachys'' meaning "short" and Latin ''rostrum'' meaning "snout" after the truncated and robust rostrum of the species. History and taxonomy ''Globidentosuchus'' was described in 2013 by T. M. Scheyer and colleagues, the ty ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Falcón
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Culebrasuchus
''Culebrasuchus'' is an extinct, monotypic genus of caiman alligatorid known from the Early to Middle Miocene (Hemingfordian) of the Panama Canal Zone of Panama. It contains a single species, ''Culebrasuchus mesoamericanus''. Discovery ''Culebrasuchus'' was first described and named by Alexander K. Hastings, Jonathan I. Bloch, Carlos A. Jaramillo, Aldo F. Rincona and Bruce J. Macfadden in 2013 based on a single holotype skull and three neck vertebrae from the Culebra Formation. ''Culebrasuchus'' is thought to be the most basal member of Caimaninae, meaning that it represents the earliest radiation of caimans in the Americas. The ancestor of ''Culebrasuchus'' likely lived farther north, perhaps in what is now southern Mexico, because before the Miocene most of Panama was underwater. The movement of ''Culebrasuchus'' into the Panama Canal Zone was an early part of the Great American Interchange in which animals dispersed between North and South America across the newly formed Is ...
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Caimaninae
A caiman (also cayman as a variant spelling) is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico, Central and South America from marshes and swamps to mangrove rivers and lakes. They have scaly skin and live a fairly nocturnal existence. They are relatively small-sized crocodilians with an average maximum weight of depending on species, with the exception of the black caiman (''Melanosuchus niger''), which can grow more than in length and weigh in excess of 1,000 kg (2,200 Ib). The black caiman is the largest caiman species in the world and is found in the slow-moving rivers and lakes that surround the Amazon basin. The smallest species is the Cuvier's dwarf caiman (''Paleosuchus palpebrosus''), which grows to long. There are six different species of caiman found throughout the watery jungle habitats of Central and Southern America. The average length for most o ...
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Alligatorinae
Alligatorinae is a subfamily within the family Alligatoridae that contains the alligators and their closest extinct relatives, and is the sister taxon to Caimaninae (the caimans). Many genera in Alligatorinae are described, but only the genus ''Alligator'' is still living, with the remaining genera extinct. Evolution Alligators and caimans split in North America during the early Tertiary or late Cretaceous (about 53 million to about 65 million years ago). The Chinese alligator split from the American alligator about 33 million years ago and likely descended from a lineage that crossed the Bering land bridge during the Neogene. The modern American alligator is well represented in the fossil record of the Pleistocene. The alligator's full mitochondrial genome was sequenced in the 1990s. The full genome, published in 2014, suggests that the alligator evolved much more slowly than mammals and birds. Phylogeny Alligatorinae is cladistically defined as ''Alligator mississippiensi ...
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Alligatoridae
The family Alligatoridae of crocodylians includes alligators, caimans and their extinct relatives. Phylogeny The superfamily Alligatoroidea includes all crocodilians (fossil and extant) that are more closely related to the American alligator than to either the Nile crocodile or the gharial. This is a stem-based definition for alligators, and is more inclusive than the crown group Alligatoridae. As a crown group, Alligatoridae only includes the last common ancestor of all extant (living) alligators, caimans, and their descendants (living or extinct), whereas Alligatoroidea, as a stem-based group, also includes more basal extinct alligator ancestors that are more closely related to living alligators than to crocodiles or gavialids. When considering only living taxa (neontology), this makes Alligatoroidea and Alligatoridae redundant. The simplified cladogram below shows Alligatoridae's relationships to other extant (living) crocodilians. The below detailed cladogram show ...
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Eocaiman
''Eocaiman'' is an extinct genus of caiman containing species living from the Early Paleocene to Miocene in what is now Argentina (Salamanca and Sarmiento Formations), Itaboraí Formation of Brazil and Colombia ( Honda Group).''Eocaiman''
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''Eocaiman'' contains three described species: ''E. cavernensis'', ''E. palaeocenicus'', and ''E. itaboraiensis'', and is typically recovered as one of the more basal members of



Necrosuchus
''Necrosuchus'' is an extinct genus of caiman from modern day Argentina that lived during the Paleocene epoch (Selandian age, about 60 million years ago). It inhabited the fluvio-lacustrine environment of the Patagonian Salamanca Formation. History and naming The fossil remains of ''Necrosuchus'' were unearthed on April 3, 1931 during the First Scarritt Expedition in the Argentinian Salamanca Formation and presented to American paleontologist Charles C. Mook. Mook however, busy with other research, was unable to describe the specimen himself. As an initial description was deemed vital to the works of several student researches at the time, Mook handed the specimen back to George G. Simpson for study. He eventually published his preliminary description in 1937, writing that his research "only carries this study as far as necessary" in the hopes of a more detailed description being later published by Mook himself. However this would not come to be and the postcranial was not describ ...
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Allognathosuchus
''Allognathosuchus'' (meaning "other jaw crocodile") is an extinct genus of alligatorine crocodylian with a complicated taxonomic history. It was named in 1921. Description ''Allognathosuchus'' was a medium-sized predator up to 1.5 m in length. This alligatorine is known for its stout jaws and bulbous teeth, found near the rear of the tooth row in upper and lower jaws. These adaptations have historically been interpreted as having been for crushing mollusks. Isolated bulbous teeth are often assigned to this genus, although such teeth are known from other crocodyliform lineages. Taxonomy The type species, ''A. polyodon'', is from the Eocene-age Bridger Formation of Wyoming; it is based on fossils that are difficult to distinguish from other Paleogene alligatorids. ''A. heterodon'' is from the Eocene-age Wasatch Formation of Wyoming, ''A. wartheni'' is from the Eocene-age Wildwood Formation (also known as the "Wildwood alligatorid" and possibly two species), and ''A. w ...
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Broad-snouted Caiman
The broad-snouted caiman (''Caiman latirostris'') is a crocodilian in the family Alligatoridae found in eastern and central South America, including southeastern Brazil, northern Argentina, Uruguay, Paraguay, and Bolivia. It is found mostly in freshwater marshes, swamps, and mangroves, usually in still or very slow-moving waters. It will often use man-made cow ponds.Britton, A. ''Caiman latirostris'' (Daudin, 1801). ''Crocodilian Species List''.http://www.flmnh.ufl.edu/cnhc/csp_clat.htm. 2009. Characteristics In the wild, adults normally grow to in length, but a few old males have been recorded to reach up to . Captive adults were found to have weighed . Most tend to be of a light olive-green color. A few individuals have spots on their faces. The most notable physical characteristic is the broad snout from which its name is derived. Borteiro et al 2009 studied the food habits of ''C. latirostris''. The snout is well adapted to rip through the dense vegetation of the marshes. Du ...
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Caiman Brevirostris
''Caiman brevirostris'' is an extinct species of caiman that lived during the Late Miocene, around 11.6 million years ago, to the end of the Miocene 5.3 million years ago in Acre and Amazonas, Brazil as well as Urumaco, Venezuela. Several specimens have been referred to the species, but only 3 of them are confidently placed in the species. ''C. brevirostris'' was originally named in 1987 on the basis of a single, incomplete rostrum with an associated mandibular ramus that had been found in Acre, Brazil. ''C. brevirostris'' is very distinct among ''Caiman'' species and caimaninae overall in that it preserves a characteristically short and robust skull that bears blunt posterior teeth that were built to break down harder foods. This was an adaption for durophagy (the diet of hard-shelled organisms), likely to crush shells of mollusks and clams which were common in the wetlands that ''C. brevirostris'' resided in. Discovery and naming ''Caiman brevirostris'' was described in 1987 ...
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Interorbital Region
The interorbital region of the skull is located between the eyes, anterior to the braincase. The form of the interorbital region may exhibit significant variation between taxonomic groups. In oryzomyine rodents, for example, the width, form, and presence of beading in the interorbital region vary among species. In birds and many other animals whose eyes are set on the side of the skull, the interorbital region normally consists of a thin interorbital septum only. This may be pierced by a hole of larger or smaller size, connecting the eye socket In anatomy, the orbit is the cavity or socket of the skull in which the eye and its appendages are situated. "Orbit" can refer to the bony socket, or it can also be used to imply the contents. In the adult human, the volume of the orbit is , of ...s. References *Weksler, M. 2006Phylogenetic relationships of oryzomyine rodents (Muroidea: Sigmodontinae): separate and combined analyses of morphological and molecular data Bulletin of the ...
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