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Granton Shrimp Bed
The Granton Shrimp Bed is a fossil-bearing deposit exposed on the southern shore of the Firth of Forth near Edinburgh, in Scotland. It is classified as a Konservat-Lagerstätten because of the exceptional quality of preservation of the fossils and is dominated by crustaceans; the deposit dates back to the Lower Carboniferous, some 359 to 323 million years ago. Location and geological setting The Granton Shrimp Bed is located on the south shore of the Firth of Forth about from the centre of Edinburgh. It consists of a layer of dolomitic limestone surrounded by Dinantian mud shales which were formed as a result of the deposit of material in either a delta plain setting or in an inter-distributary bay, where sedimentation occurred because of flood-generated incursions. The shrimp bed resulted from the periodic inundations of marine water into stagnant lagoons, each incursion leaving a lamina of limestone rich in the fossils of soft-bodied marine invertebrates and other animals. T ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the ...
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Polychaete
Polychaeta () is a paraphyletic class (biology), class of generally marine invertebrate, marine annelid worms, common name, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin. More than 10,000 species are described in this class. Common representatives include the lugworm (''Arenicola marina'') and the Alitta virens, sandworm or Alitta succinea, clam worm ''Alitta''. Polychaetes as a class are robust and widespread, with species that live in the coldest ocean temperatures of the abyssal plain, to forms which tolerate the extremely high temperatures near hydrothermal vents. Polychaetes occur throughout the Earth's oceans at all depths, from forms that live as plankton near the surface, to a 2- to 3-cm specimen (still unclassified) observed by the robot ocean probe Nereus (underwater vehicle), ''Nereus'' at the bottom of the Challenger Deep, the deepes ...
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Carboniferous Paleontological Sites
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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Paleontology In Scotland
Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (geology), epoch (roughly 11,700 years before present). It includes the study of fossils to classify organisms and study their biological interaction, interactions with each other and their environments (their paleoecology). Paleontological observations have been documented as far back as the 5th century BC. The science became established in the 18th century as a result of Georges Cuvier's work on comparative anatomy, and developed rapidly in the 19th century. The term itself originates from ancient language, Greek (, "old, ancient"), (, (Genitive, gen. ), "being, creature"), and (, "speech, thought, study"). Paleontology lies on the border between biology and geology, but differs from archaeology in that it excludes the study of anatomically modern humans. It now uses techniques drawn from a wide range ...
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Bioturbation
Bioturbation is defined as the reworking of soils and sediments by animals or plants. It includes burrowing, ingestion, and defecation of sediment grains. Bioturbating activities have a profound effect on the environment and are thought to be a primary driver of biodiversity. The formal study of bioturbation began in the 1800s by Charles Darwin experimenting in his garden. The disruption of aquatic sediments and terrestrial soils through bioturbating activities provides significant ecosystem services. These include the alteration of nutrients in aquatic sediment and overlying water, shelter to other species in the form of burrows in terrestrial and water ecosystems, and soil production on land.Shaler, N. S., 1891, The origin and nature of soils, in Powell, J. W., ed., USGS 12th Annual report 1890-1891: Washington, D.C., Government Printing Office, p. 213-45. Bioturbators are deemed ecosystem engineers because they alter resource availability to other species through the physical ch ...
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Geology Today
''Geology Today'' is a bimonthly peer-reviewed scientific journal published by Wiley on behalf of the Geologists' Association and the Geological Society of London. The editor-in-chief is Peter Doyle (Geological Society of London). The journal covers all aspects of the Earth sciences. Abstracting and indexing The journal is abstracted and indexed in Aquatic Sciences and Fisheries Abstracts, EBSCO databases, ProQuest databases, Scopus, and The Zoological Record ''The Zoological Record'' (''ZR'') is an electronic index of zoological literature that also serves as the unofficial register of scientific names in zoology. It was started as a print publication in 1864 by the Zoological Society of London, a .... References External links * Geology journals Wiley (publisher) academic journals Geological Society of London academic journals Publications established in 1985 Geologists' Association 1985 establishments in the United Kingdom {{geology-journal-stub ...
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Royal Scottish Museum
The National Museum of Scotland in Edinburgh, Scotland, was formed in 2006 with the merger of the new Museum of Scotland, with collections relating to Scottish antiquities, culture and history, and the adjacent Royal Scottish Museum (opened in 1866 as the Edinburgh Museum of Science and Art, renamed in 1904, and for the period between 1985 and the merger named the Royal Museum of Scotland or simply the Royal Museum), with international collections covering science and technology, natural history, and world cultures. The two connected buildings stand beside each other on Chambers Street, by the intersection with the George IV Bridge, in central Edinburgh. The museum is part of National Museums Scotland. Admission is free. The two buildings retain distinctive characters: the Museum of Scotland is housed in a modern building opened in 1998, while the former Royal Museum building was begun in 1861 and partially opened in 1866, with a Victorian Venetian Renaissance facade and a gr ...
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Waterstonella
''Waterstonella grantonensis'' is a species of fossil crustacean so distinct from other crustaceans that it has been placed in its own genus, ''Waterstonella'', family (biology), family, Waterstonellidae, and Order (biology), order, Waterstonellidea. It is named after Charles Waterstone, keeper of geology at the Royal Scottish Museum, while the specific name (zoology), specific epithet commemorates the location where the fossil was found, the Granton shrimp beds, near Edinburgh. References

Prehistoric crustacean genera Carboniferous crustaceans Prehistoric life of Europe {{paleo-crustacean-stub ...
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Frederick Schram
Frederick Robert Schram (born August 11, 1943, in Chicago, Illinois, Chicago, Illinois) is an Americans, American palaeontologist and carcinologist. He received his Bachelor of Science, B.S. in biology from Loyola University Chicago in 1965, and a Doctor of Philosophy, Ph.D. on palaeozoology from the University of Chicago in 1968 . He has written over 200 papers on various aspects of crustacean biology, Taxonomy (biology), taxonomy and systematics, as well as several books, including the standard text ''Crustacea'' . In 1983, he founded the journal ''Crustacean Issues'', which he continued to edit for over twenty years. Much of his career has been spent at the Universiteit van Amsterdam, the Netherlands, from which he retired in 2005. In July 2005, he became the Editor of the ''Journal of Crustacean Biology'' . References

American carcinologists American paleontologists 1943 births Living people Scientists from Chicago American expatriates in the Netherla ...
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Tealliocaris
''Tealliocaris'' is an extinct genus of pygocephalomorphans from the Carboniferous. Species The genus contains eight described species: *''Tealliocaris caudafimbriata'' *''Tealliocaris etheridgei'' *''Tealliocaris formosa'' *''Tealliocaris loudonensis'' *''Tealliocaris robusta'' *''Tealliocaris tarrassiana'' *''Tealliocaris walloniensis'' *''Tealliocaris woodwardi'' References

* Fossils (Smithsonian Handbooks) by David Ward (Page 69) Prehistoric Malacostraca Prehistoric crustacean genera Carboniferous crustaceans Fossils of Great Britain Fossil taxa described in 1908 {{Malacostraca-stub ...
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Conodont
Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which are usually found in isolation and are now called conodont elements. Knowledge about soft tissues remains limited. They existed in the world's oceans for over 300 million years, from the Cambrian to the beginning of the Jurassic. Conodont elements are widely used as index fossils, fossils used to define and identify geological periods. The animals are also called Conodontophora (conodont bearers) to avoid ambiguity. Discovery and understanding of conodonts The teeth-like fossils of the conodont were first discovered by Heinz Christian Pander and the results published in Saint Petersburg, Russia, in 1856. The name ''pander'' is commonly used in scientific names of conodonts. It was only in the early 1980s that the first fossil evidence of ...
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