Founder Takes All
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Founder Takes All
The Founder Takes All (FTA) hypothesis refers to the evolutionary advantages conferred to first-arriving lineages in an ecosystem. Overview Density-dependent processes such as gene surfing, high-density blocking, and competitive exclusion can play crucial roles in the spatial structuring of biodiversity. These interrelated demographic processes can generate striking geographic contrasts in the distributions of genes and species. Density-dependent processes refer to biogeographical processes in which population density (relative to resources) constrains the ability of new arriving lineages to establish It is proposed that well-studied evolutionary and ecological biogeographic patterns of postglacial recolonization, progressive island colonization, microbial sectoring, and even the "Out of Africa" pattern of human expansion are fundamentally similar. All these patterns are underpinned by the "founder takes all" density-dependent principle. For example, following a large-scale ea ...
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Phylogeography
Phylogeography is the study of the historical processes that may be responsible for the past to present geographic distributions of genealogical lineages. This is accomplished by considering the geographic distribution of individuals in light of genetics, particularly population genetics. This term was introduced to describe geographically structured genetic signals within and among species. An explicit focus on a species' biogeography/biogeographical past sets phylogeography apart from classical population genetics and phylogenetics. Past events that can be inferred include population expansion, population bottlenecks, vicariance, dispersal, and migration. Recently developed approaches integrating coalescent theory or the genealogical history of alleles and distributional information can more accurately address the relative roles of these different historical forces in shaping current patterns. Development The term phylogeography was first used by John Avise in his 1987 work '' ...
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Genetic Drift
Genetic drift, also known as allelic drift or the Wright effect, is the change in the frequency of an existing gene variant (allele) in a population due to random chance. Genetic drift may cause gene variants to disappear completely and thereby reduce genetic variation. It can also cause initially rare alleles to become much more frequent and even fixed. When few copies of an allele exist, the effect of genetic drift is more notable, and when many copies exist, the effect is less notable. In the middle of the 20th century, vigorous debates occurred over the relative importance of natural selection versus neutral processes, including genetic drift. Ronald Fisher, who explained natural selection using Mendelian genetics, held the view that genetic drift plays at most a minor role in evolution, and this remained the dominant view for several decades. In 1968, population geneticist Motoo Kimura rekindled the debate with his neutral theory of molecular evolution, which claims that ...
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Peripatric Speciation
Peripatric speciation is a mode of speciation in which a new species is formed from an isolated peripheral population. Since peripatric speciation resembles allopatric speciation, in that populations are isolated and prevented from exchanging genes, it can often be difficult to distinguish between them. Nevertheless, the primary characteristic of peripatric speciation proposes that one of the populations is much smaller than the other. The terms peripatric and peripatry are often used in biogeography, referring to organisms whose ranges are closely adjacent but do not overlap, being separated where these organisms do not occur—for example on an oceanic island compared to the mainland. Such organisms are usually closely related (''e.g.'' sister species); their distribution being the result of peripatric speciation. The concept of peripatric speciation was first outlined by the evolutionary biologist Ernst Mayr in 1954.Ernst Mayr. (1954). Change of genetic environment and evolut ...
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Founder Effect
In population genetics, the founder effect is the loss of genetic variation that occurs when a new population is established by a very small number of individuals from a larger population. It was first fully outlined by Ernst Mayr in 1942, using existing theoretical work by those such as Sewall Wright. As a result of the loss of genetic variation, the new population may be distinctively different, both genotypically and phenotypically, from the parent population from which it is derived. In extreme cases, the founder effect is thought to lead to the speciation and subsequent evolution of new species. In the figure shown, the original population has nearly equal numbers of blue and red individuals. The three smaller founder populations show that one or the other color may predominate (founder effect), due to random sampling of the original population. A population bottleneck may also cause a founder effect, though it is not strictly a new population. The founder effect occurs wh ...
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Invasive Species
An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native species that become harmful to their native environment after human alterations to its food webfor example the purple sea urchin (''Strongylocentrotus purpuratus'') which has decimated kelp forests along the northern California coast due to overharvesting of its natural predator, the California sea otter (''Enhydra lutris''). Since the 20th century, invasive species have become a serious economic, social, and environmental threat. Invasion of long-established ecosystems by organisms is a natural phenomenon, but human-facilitated introductions have greatly increased the rate, scale, and geographic range of ...
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Gene Flow
In population genetics, gene flow (also known as gene migration or geneflow and allele flow) is the transfer of genetic material from one population to another. If the rate of gene flow is high enough, then two populations will have equivalent allele frequencies and therefore can be considered a single effective population. It has been shown that it takes only "one migrant per generation" to prevent populations from diverging due to drift. Populations can diverge due to selection even when they are exchanging alleles, if the selection pressure is strong enough. Gene flow is an important mechanism for transferring genetic diversity among populations. Migrants change the distribution of genetic diversity among populations, by modifying allele frequencies (the proportion of members carrying a particular variant of a gene). High rates of gene flow can reduce the genetic differentiation between the two groups, increasing homogeneity. For this reason, gene flow has been thought to con ...
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Extirpation
Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinctions. Local extinctions mark a change in the ecology of an area. In recent times, local extinction has sometimes been followed by a replacement of the species taken from other locations; wolf reintroduction is an example of this. The term "local extinction" is highly vernacular. The more proper biological term is ''extirpation''. Discussion Glaciation can lead to local extinction. This was the case during the Pleistocene glaciation event in North America. During this period, most of the native North American species of earthworm were killed in places covered by glaciation. This left them open for colonization by European earthworms brought over in soil from Europe. Species naturally become extirpated from islands over time. The number ...
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Deglaciation
Deglaciation is the transition from full glacial conditions during ice ages, to warm interglacials, characterized by global warming and sea level rise due to change in continental ice volume. Thus, it refers to the retreat of a glacier, an ice sheet or frozen surface layer, and the resulting exposure of the Earth's surface. The decline of the cryosphere due to ablation can occur on any scale from global to localized to a particular glacier. After the Last Glacial Maximum (ca. 21,000 years ago), the last deglaciation begun, which lasted until the early Holocene. Around much of Earth, deglaciation during the last 100 years has been accelerating as a result of climate change, partly brought on by anthropogenic changes to greenhouse gases. The previous deglaciation took place from approximately 22  ka until 11.5 ka. This occurred when there was an annual mean atmospheric temperature on the earth that increased by roughly 5 °C, which was also accompanied by regional hi ...
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Priority Effect
In ecology, a priority effect is an impact that a particular species can have on community development due to prior arrival at a site. There are two basic types: An ''inhibitory priority effect'' which occurs when a species that arrives first at a site negatively impacts a species that arrives later by reducing the availability of space or resources. A ''facilitative priority effect'' occurs when a species that arrives first at a site alters abiotic or biotic conditions in ways that positively impact a species arriving later. Priority effects are a central and pervasive element of ecological community development which have important implications for natural systems as well as ecological restoration efforts. Theoretical foundation Community succession theory Early in the 20th century, Frederic Clements and other plant ecologists suggested that ecological communities develop in a linear, directional manner towards a final, stable end-point: the climax community. Clements indicate ...
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Hypothesis
A hypothesis (plural hypotheses) is a proposed explanation for a phenomenon. For a hypothesis to be a scientific hypothesis, the scientific method requires that one can test it. Scientists generally base scientific hypotheses on previous observations that cannot satisfactorily be explained with the available scientific theories. Even though the words "hypothesis" and "theory" are often used interchangeably, a scientific hypothesis is not the same as a scientific theory. A working hypothesis is a provisionally accepted hypothesis proposed for further research in a process beginning with an educated guess or thought. A different meaning of the term ''hypothesis'' is used in formal logic, to denote the antecedent of a proposition; thus in the proposition "If ''P'', then ''Q''", ''P'' denotes the hypothesis (or antecedent); ''Q'' can be called a consequent. ''P'' is the assumption in a (possibly counterfactual) ''What If'' question. The adjective ''hypothetical'', meaning "hav ...
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