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Flehmen Response
The flehmen response (; from German ''flehmen'', to bare the upper teeth, and Upper Saxon German ''flemmen'', to look spiteful), also called the flehmen position, flehmen reaction, flehmen grimace, flehming, or flehmening, is a behavior in which an animal curls back its upper lip exposing its front teeth, inhales with the nostrils usually closed, and then often holds this position for several seconds. It may be performed over a sight or substance of particular interest to the animal, or may be performed with the neck stretched and the head held high in the air. Flehmen is performed by a wide range of mammals, including ungulates and felids. The behavior facilitates the transfer of pheromones and other scents into the vomeronasal organ (VNO, or Jacobson's organ) located above the roof of the mouth via a duct which exits just behind the front teeth of the animal. Etymology The word originates from the German verb ''flehmen'', to bare the upper teeth. It comes from the Upper Saxon ...
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Animal
Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a bilaterally symmetric body plan. The Bilateria include the protostomes, containing animals such as nematodes, arthropods, flatworms, annelids and molluscs, and the deuterostomes, containing t ...
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Reproductive Synchrony
''Reproductive synchrony'' is a term used in evolutionary biology and behavioral ecology. Reproductive synchrony—sometimes termed "ovulatory synchrony"—may manifest itself as "breeding seasonality". Where females undergo regular menstruation, "menstrual synchrony" is another possible term. Reproduction is said to be synchronised when fertile matings across a population are temporarily clustered, resulting in multiple conceptions (and consequent births) within a restricted time window. In marine and other aquatic contexts, the phenomenon may be referred to as mass spawning. Mass spawning has been observed and recorded in a large number of phyla, including in coral communities within the Great Barrier Reef. In primates, reproductive synchrony usually takes the form of conception and birth seasonality. The regulatory "clock", in this case, is the sun's position in relation to the tilt of the earth. In nocturnal or partly nocturnal primates—for example, owl monkeys—the peri ...
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Estrus
The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous phases, otherwise known as "rest" phases, or by pregnancies. Typically, estrous cycles repeat until death. These cycles are widely variable in duration and frequency depending on the species.Bronson, F. H., 1989. Mammalian Reproductive Biology. University of Chicago Press, Chicago, IL, USA. Some animals may display bloody vaginal discharge, often mistaken for menstruation. Many mammals used in commercial agriculture, such as cattle and sheep, may have their estrous cycles artificially controlled with hormonal medications for optimum productivity. The male equivalent, seen primarily in ruminants, is called rut. Differences from the menstrual cycle Mammals share the same reproductive system, including the regulatory hypothalamic syste ...
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Sheep Reproduction
Domestic sheep reproduce sexually much like other mammals, and their reproductive strategy is furthermore very similar to other domestic herd animals. A flock of sheep is generally mated by a single ram, which has either been chosen by a farmer or has established dominance through physical contest with other rams (in feral populations). Most sheep have a breeding season (''tupping'') in the autumn, though some are able to breed year-round. Largely as a result of the influence of humans in sheep breeding, ewes often produce multiple lambs. This increase in the lamb births, both in number and birth weight, may cause problems in delivery and lamb survival, requiring the intervention of shepherds. Sexual behavior Ewes generally reach sexual maturity at six to eight months of age, and rams generally at four to six (ram lambs have occasionally been known to impregnate their mothers at two months). Sheep are seasonally polyoestrus animals. Ewes enter into oestrus cycles about every 17 ...
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Pheromones
A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavior of the receiving individuals. There are ''alarm pheromones'', ''food trail pheromones'', ''sex pheromones'', and many others that affect behavior or physiology. Pheromones are used by many organisms, from basic unicellular prokaryotes to complex multicellular eukaryotes. Their use among insects has been particularly well documented. In addition, some vertebrates, plants and ciliates communicate by using pheromones. The ecological functions and evolution of pheromones are a major topic of research in the field of chemical ecology. Background The portmanteau word "pheromone" was coined by Peter Karlson and Martin Lüscher in 1959, based on the Greek φερω ''pheroo'' ('I carry') and ὁρμων ''hormon'' ('stimulating'). Pheromones ...
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Organ (anatomy)
In biology, an organ is a collection of tissues joined in a structural unit to serve a common function. In the hierarchy of life, an organ lies between tissue and an organ system. Tissues are formed from same type cells to act together in a function. Tissues of different types combine to form an organ which has a specific function. The intestinal wall for example is formed by epithelial tissue and smooth muscle tissue. Two or more organs working together in the execution of a specific body function form an organ system, also called a biological system or body system. An organ's tissues can be broadly categorized as parenchyma, the functional tissue, and stroma, the structural tissue with supportive, connective, or ancillary functions. For example, the gland's tissue that makes the hormones is the parenchyma, whereas the stroma includes the nerves that innervate the parenchyma, the blood vessels that oxygenate and nourish it and carry away its metabolic wastes, and the con ...
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Chemosensory
A chemoreceptor, also known as chemosensor, is a specialized sensory receptor which transduces a chemical substance ( endogenous or induced) to generate a biological signal. This signal may be in the form of an action potential, if the chemoreceptor is a neuron, or in the form of a neurotransmitter that can activate a nerve fiber if the chemoreceptor is a specialized cell, such as taste receptors, or an internal peripheral chemoreceptor, such as the carotid bodies. In physiology, a chemoreceptor detects changes in the normal environment, such as an increase in blood levels of carbon dioxide (hypercapnia) or a decrease in blood levels of oxygen (hypoxia), and transmits that information to the central nervous system which engages body responses to restore homeostasis. In bacteria, chemoreceptors are essential in the mediation of chemotaxis. Cellular chemoreceptors In prokaryotes Bacteria utilize complex long helical proteins as chemoreceptors, permitting signals to trave ...
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Volatile Organic Compounds
Volatile organic compounds (VOCs) are organic compounds that have a high vapour pressure at room temperature. High vapor pressure correlates with a low boiling point, which relates to the number of the sample's molecules in the surrounding air, a trait known as volatility. VOCs are responsible for the odor of scents and perfumes as well as pollutants. VOCs play an important role in communication between animals and plants, e.g. attractants for pollinators, protection from predation, and even inter-plant interactions. Some VOCs are dangerous to human health or cause harm to the environment. Anthropogenic VOCs are regulated by law, especially indoors, where concentrations are the highest. Most VOCs are not acutely toxic, but may have long-term chronic health effects. Definitions Diverse definitions of the term VOC are in use. Canada Health Canada classifies VOCs as organic compounds that have boiling points roughly in the range of . The emphasis is placed on commonly encountered VO ...
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Organic Compound
In chemistry, organic compounds are generally any chemical compounds that contain carbon-hydrogen or carbon-carbon bonds. Due to carbon's ability to catenate (form chains with other carbon atoms), millions of organic compounds are known. The study of the properties, reactions, and syntheses of organic compounds comprise the discipline known as organic chemistry. For historical reasons, a few classes of carbon-containing compounds (e.g., carbonate salts and cyanide salts), along with a few other exceptions (e.g., carbon dioxide, hydrogen cyanide), are not classified as organic compounds and are considered inorganic. Other than those just named, little consensus exists among chemists on precisely which carbon-containing compounds are excluded, making any rigorous definition of an organic compound elusive. Although organic compounds make up only a small percentage of Earth's crust, they are of central importance because all known life is based on organic compounds. Li ...
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Incisive Foramen
In the human mouth, the incisive foramen (also known as: "''anterior palatine foramen''", or "''nasopalatine foramen''") is the opening of the incisive canals on the hard palate immediately behind the incisor teeth. It gives passage to blood vessels and nerves. The incisive foramen is situated within the incisive fossa of the maxilla. The incisive foramen is used as an anatomical landmark for defining the severity of cleft lip and cleft palate. The incisive foramen exists in a variety of species. Structure The incisive foramen is a funnel-shaped opening of the in the bone of the oral hard palate representing the inferior termination of the incisive canal. An oral prominence - the incisive papilla - overlies the incisive fossa. The incisive foramen is situated immediately behind the incisor teeth, and in between the two premaxillae. Contents The incisive foramen allows for blood vessels and nerves to pass. These include: * the pterygopalatine nerves to the hard palate. ...
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Incisive Papilla
The incisive papilla is an oval midline mucosal prominence of the anterior hard palate overlying the incisive fossa. It is situated posteriorly to the central incisors, and represents the anterior extremity of the palatine raphe. The incisive papilla marks the position of the foetal nasopalatine canal. Anatomy Variation Though generally referred to as round or oval in shape, the shape of the incisive papilla has been noted to vary in shape, its other potential shapes have been described as including: pear, spindle, flame, cylindrical, tapering, or dumb bell shaped. Microanatomy The incisive papilla consist of dense connective tissue. It is lined with simple or pseudostratified columnar epithelium, and is often keratinized. Clinical significance Pressure exerted upon the incisive papilla by maxillary dentures may cause pain or discomfort. The incisive papilla is utilized as an anatomical landmark when administering a nasopalatine nerve block Nerve block or regional ...
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