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Flehmen
The flehmen response (; from German ''flehmen'', to bare the upper teeth, and Upper Saxon German ''flemmen'', to look spiteful), also called the flehmen position, flehmen reaction, flehmen grimace, flehming, or flehmening, is a behavior in which an animal curls back its upper lip exposing its front teeth, inhales with the nostrils usually closed, and then often holds this position for several seconds. It may be performed over a sight or substance of particular interest to the animal, or may be performed with the neck stretched and the head held high in the air. Flehmen is performed by a wide range of mammals, including ungulates and felids. The behavior facilitates the transfer of pheromones and other scents into the vomeronasal organ (VNO, or Jacobson's organ) located above the roof of the mouth via a duct which exits just behind the front teeth of the animal. Etymology The word originates from the German verb ''flehmen'', to bare the upper teeth. It comes from the Upper Saxon ...
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Vomeronasal Organ
The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods. The name is derived from the fact that it lies adjacent to the unpaired vomer bone (from Latin 'plowshare', for its shape) in the nasal septum. It is present and functional in all snakes and lizards, and in many mammals, including cats, dogs, cattle, pigs, and some primates. Some humans may have physical remnants of a VNO, but it is vestigial and non-functional. The VNO contains the cell bodies of sensory neurons which have receptors that detect specific non-volatile (liquid) organic compounds which are conveyed to them from the environment. These compounds emanate from prey, predators, and the compounds called sex pheromones from potential mates. Activation of the VNO triggers an appropriate behavioral response to the presence of ...
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Jacobson's Organ
The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods. The name is derived from the fact that it lies adjacent to the unpaired vomer bone (from Latin 'plowshare', for its shape) in the nasal septum. It is present and functional in all snakes and lizards, and in many mammals, including cats, dogs, cattle, pigs, and some primates. Some humans may have physical remnants of a VNO, but it is vestigial and non-functional. The VNO contains the cell bodies of sensory neurons which have receptors that detect specific non-volatile (liquid) organic compounds which are conveyed to them from the environment. These compounds emanate from prey, predators, and the compounds called sex pheromones from potential mates. Activation of the VNO triggers an appropriate behavioral response to the presence of one o ...
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Karl Max Schneider
Karl Max Schneider (13 March 1887 - 26 October 1955) was an East German zoologist who served as director for the Leipzig Zoo from 1934 till his death. He was an expert on lions and their biology and was responsible for coining the behavioural term " flehmen". Biography Schneider was born in Callenberg, Lichtenstein, one of six siblings in a merchant family. He studied in his hometown and then at Waldenberg where he became a teacher from 1908 working in Meerane. In 1912 he received an Abitur from the Realgymnasium Freiberg and a degree in science in 1910 from the University of Leipzig. In 1913 he wrote a thesis on the philosophy of Heinrich Rickert's transcendentalism Transcendentalism is a philosophical movement that developed in the late 1820s and 1830s in New England. "Transcendentalism is an American literary, political, and philosophical movement of the early nineteenth century, centered around Ralph Wald ..., but due to the onset of World War I he received a doctorate only ...
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Sense Organ
A sense is a biological system used by an organism for sensation, the process of gathering information about the world through the detection of Stimulus (physiology), stimuli. (For example, in the human body, the brain which is part of the central nervous system receives signals from the senses which continuously receive information from the environment, interprets these signals, and causes the body to respond, either chemically or physically.) Although traditionally five human senses were identified as such (namely Visual perception, sight, Olfaction, smell, Somatosensory system, touch, taste, and hearing), it is now recognized that there are many more. Senses used by non-human organisms are even greater in variety and number. During sensation, sense organs collect various stimuli (such as a sound or smell) for Transduction (physiology), transduction, meaning transformation into a form that can be understood by the brain. Sensation and perception are fundamental to nearly every ...
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Olfactory
The sense of smell, or olfaction, is the special sense through which smells (or odors) are perceived. The sense of smell has many functions, including detecting desirable foods, hazards, and pheromones, and plays a role in taste. In humans, it occurs when an odor binds to a receptor within the nasal cavity, transmitting a signal through the olfactory system. Glomeruli aggregate signals from these receptors and transmit them to the olfactory bulb, where the sensory input will start to interact with parts of the brain responsible for smell identification, memory, and emotion. There are many different causes for alteration, lack, or disturbance to a normal sense of smell, and can include damage to the nose or smell receptors, or central problems affecting the brain. Some causes include upper respiratory infections, traumatic brain injury, and neurodegenerative disease. History of study Early scientific study of the sense of smell includes the extensive doctoral dissertation of ...
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Felid
Felidae () is the family of mammals in the order Carnivora colloquially referred to as cats, and constitutes a clade. A member of this family is also called a felid (). The term "cat" refers both to felids in general and specifically to the domestic cat ('' Felis catus''). Felidae species exhibit the most diverse fur pattern of all terrestrial carnivores. Cats have retractile claws, slender muscular bodies and strong flexible forelimbs. Their teeth and facial muscles allow for a powerful bite. They are all obligate carnivores, and most are solitary predators ambushing or stalking their prey. Wild cats occur in Africa, Europe, Asia and the Americas. Some wild cat species are adapted to forest habitats, some to arid environments, and a few also to wetlands and mountainous terrain. Their activity patterns range from nocturnal and crepuscular to diurnal, depending on their preferred prey species. Reginald Innes Pocock divided the extant Felidae into three subfamilies: the Panth ...
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Incisive Papilla
The incisive papilla is an oval midline mucosal prominence of the anterior hard palate overlying the incisive fossa. It is situated posteriorly to the central incisors, and represents the anterior extremity of the palatine raphe. The incisive papilla marks the position of the foetal nasopalatine canal. Anatomy Variation Though generally referred to as round or oval in shape, the shape of the incisive papilla has been noted to vary in shape, its other potential shapes have been described as including: pear, spindle, flame, cylindrical, tapering, or dumb bell shaped. Microanatomy The incisive papilla consist of dense connective tissue. It is lined with simple or pseudostratified columnar epithelium, and is often keratinized. Clinical significance Pressure exerted upon the incisive papilla by maxillary dentures may cause pain or discomfort. The incisive papilla is utilized as an anatomical landmark when administering a nasopalatine nerve block; the needle is inserted at a ...
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Incisive Foramen
In the human mouth, the incisive foramen (also known as: "''anterior palatine foramen''", or "''nasopalatine foramen''") is the opening of the incisive canals on the hard palate immediately behind the incisor teeth. It gives passage to blood vessels and nerves. The incisive foramen is situated within the incisive fossa of the maxilla. The incisive foramen is used as an anatomical landmark for defining the severity of cleft lip and cleft palate. The incisive foramen exists in a variety of species. Structure The incisive foramen is a funnel-shaped opening of the in the bone of the oral hard palate representing the inferior termination of the incisive canal. An oral prominence - the incisive papilla - overlies the incisive fossa. The incisive foramen is situated immediately behind the incisor teeth, and in between the two premaxillae. Contents The incisive foramen allows for blood vessels and nerves to pass. These include: * the pterygopalatine nerves to the hard palate. * ...
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Organic Compound
In chemistry, organic compounds are generally any chemical compounds that contain carbon-hydrogen or carbon-carbon bonds. Due to carbon's ability to catenate (form chains with other carbon atoms), millions of organic compounds are known. The study of the properties, reactions, and syntheses of organic compounds comprise the discipline known as organic chemistry. For historical reasons, a few classes of carbon-containing compounds (e.g., carbonate salts and cyanide salts), along with a few other exceptions (e.g., carbon dioxide, hydrogen cyanide), are not classified as organic compounds and are considered inorganic. Other than those just named, little consensus exists among chemists on precisely which carbon-containing compounds are excluded, making any rigorous definition of an organic compound elusive. Although organic compounds make up only a small percentage of Earth's crust, they are of central importance because all known life is based on organic compounds. Living t ...
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Volatile Organic Compounds
Volatile organic compounds (VOCs) are organic compounds that have a high vapour pressure at room temperature Colloquially, "room temperature" is a range of air temperatures that most people prefer for indoor settings. It feels comfortable to a person when they are wearing typical indoor clothing. Human comfort can extend beyond this range depending on .... High vapor pressure correlates with a low boiling point, which relates to the number of the sample's molecules in the surrounding air, a trait known as volatility (chemistry), volatility. VOCs are responsible for the odor of scents and perfumes as well as pollutants. VOCs play an important role in communication between animals and plants, e.g. attractants for pollinators, protection from predation, and even inter-plant interactions. Some VOCs are dangerous to human health or cause harm to the natural environment, environment. Human impact on the environment, Anthropogenic VOCs are regulated by law, especially indoors, where c ...
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Chemosensory
A chemoreceptor, also known as chemosensor, is a specialized sensory receptor which transduces a chemical substance ( endogenous or induced) to generate a biological signal. This signal may be in the form of an action potential, if the chemoreceptor is a neuron, or in the form of a neurotransmitter that can activate a nerve fiber if the chemoreceptor is a specialized cell, such as taste receptors, or an internal peripheral chemoreceptor, such as the carotid bodies. In physiology, a chemoreceptor detects changes in the normal environment, such as an increase in blood levels of carbon dioxide (hypercapnia) or a decrease in blood levels of oxygen (hypoxia), and transmits that information to the central nervous system which engages body responses to restore homeostasis. In bacteria, chemoreceptors are essential in the mediation of chemotaxis. Cellular chemoreceptors In prokaryotes Bacteria utilize complex long helical proteins as chemoreceptors, permitting signals to travel lo ...
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Nasal Bone
The nasal bones are two small oblong bones, varying in size and form in different individuals; they are placed side by side at the middle and upper part of the face and by their junction, form the bridge of the upper one third of the nose. Each has two surfaces and four borders. Structure The two nasal bones are joined at the midline internasal suture and make up the bridge of the nose. Surfaces The ''outer surface'' is concavo-convex from above downward, convex from side to side; it is covered by the procerus and nasalis muscles, and perforated about its center by a foramen, for the transmission of a small vein. The ''inner surface'' is concave from side to side, and is traversed from above downward, by a groove for the passage of a branch of the nasociliary nerve. Articulations The nasal articulates with four bones: two of the cranium, the frontal and ethmoid, and two of the face, the opposite nasal and the maxilla. Other animals In primitive bony fish and tetrapod ...
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