Evolution Of Color Vision In Primates
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Evolution Of Color Vision In Primates
The evolution of color vision in primates is highly unusual compared to most eutherian mammals. A remote vertebrate ancestor of primates possessed tetrachromacy, but nocturnal, warm-blooded, mammalian ancestors lost two of four cones in the retina at the time of dinosaurs. Most teleost fish, reptiles and birds are therefore tetrachromatic while most mammals are strictly dichromats, the exceptions being some primates and marsupials, who are trichromats, and many marine mammals, who are monochromats. Cones and opsins While color vision is dependent on many factors, discussion of the evolution of color vision is typically simplified to two factors: * the breadth of the visible spectrum (which wavelengths of light can be detected), and * the dimensionality of the color gamut (e.g. dichromacy vs. tetrachromacy). In vertebrates, both of these are almost perfectly correlated to an individual's cone complement. The retina comprises several different classes of photoreceptors, i ...
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Olive Baboon Papio Anubis In Tanzania 4426 Cropped Nevit
The olive, botanical name ''Olea europaea'', meaning 'European olive' in Latin, is a species of small tree or shrub in the family Oleaceae, found traditionally in the Mediterranean Basin. When in shrub form, it is known as ''Olea europaea'' 'Montra', dwarf olive, or little olive. The species is cultivated in all the countries of the Mediterranean, as well as in Australia, New Zealand, North and South America and South Africa. ''Olea europaea'' is the type species for the genus ''Olea''. The olive's fruit, also called an "olive", is of major agricultural importance in the Mediterranean region as the source of olive oil; it is one of the core ingredients in Mediterranean cuisine. The tree and its fruit give their name to the plant family, which also includes species such as lilac, jasmine, forsythia, and the true ash tree. Thousands of cultivars of the olive tree are known. Olive cultivars may be used primarily for oil, eating, or both. Olives cultivated for consumption are ge ...
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Marine Mammal
Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as seals, whales, manatees, sea otters and polar bears. They are an informal group, unified only by their reliance on marine environments for feeding and survival. Marine mammal adaptation to an aquatic lifestyle varies considerably between species. Both cetaceans and sirenians are fully aquatic and therefore are obligate water dwellers. Seals and sea-lions are semiaquatic; they spend the majority of their time in the water but need to return to land for important activities such as mating, breeding and molting. In contrast, both otters and the polar bear are much less adapted to aquatic living. The diets of marine mammals vary considerably as well; some eat zooplankton, others eat fish, squid, shellfish, or sea-grass, and a few eat other mammals. While the number of marine mammals is small compared to those found on land, their roles in various ...
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Howler Monkey
Howler monkeys (genus ''Alouatta'', monotypic in subfamily Alouattinae) are the most widespread primate genus in the Neotropics and are among the largest of the platyrrhines along with the muriquis (''Brachyteles''), the spider monkeys (''Ateles'') and woolly monkeys (''Lagotrix''). These monkeys are native to South and Central American forests. They are famous for their loud howls, which can travel more than a mile through dense rain forest. Fifteen species are recognized. Previously classified in the family Cebidae, they are now placed in the family Atelidae. They are primarily folivores but also significant frugivores, acting as seed dispersal agents through their digestive system and their locomotion. Threats include human predation, habitat destruction, and capture for pets or zoo animals. Classification * ''A. palliata'' group ** Coiba Island howler, ''Alouatta coibensis'' *** ''Alouatta coibensis coibensis'' *** Azuero howler, ''Alouatta coibensis trabeata'' ** Mantled h ...
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New World Monkey
New World monkeys are the five families of primates that are found in the tropical regions of Mexico, Central and South America: Callitrichidae, Cebidae, Aotidae, Pitheciidae, and Atelidae. The five families are ranked together as the Ceboidea (), the only extant superfamily in the parvorder Platyrrhini (). Platyrrhini is derived from the Greek for "broad nosed", and their noses are flatter than those of other simians, with sideways-facing nostrils. Monkeys in the family Atelidae, such as the spider monkey, are the only primates to have prehensile tails. New World monkeys' closest relatives are the other simians, the Catarrhini ("down-nosed"), comprising Old World monkeys and apes. New World monkeys descend from African simians that colonized South America, a line that split off about 40 million years ago. Evolutionary history About 40 million years ago, the Simiiformes infraorder split into the parvorders Platyrrhini (New World monkeys) and Catarrhini (apes and Old World mon ...
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Old World Monkey
Old World monkey is the common English name for a family of primates known taxonomically as the Cercopithecidae (). Twenty-four genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons (genus '' Papio''), red colobus (genus '' Piliocolobus'') and macaques (genus '' Macaca''). Common names for other Old World monkeys include the talapoin, guenon, colobus, douc (douc langur, genus ''Pygathrix''), vervet, gelada, mangabey (a group of genera), langur, mandrill, surili (''Presbytis''), patas, and proboscis monkey. Phylogenetically, they are more closely related to apes than to New World monkeys. They diverged from a common ancestor of New World monkeys around 45 to 55 million years ago. The smallest Old World monkey is the talapoin, with a head and body 34–37 cm in length, and weighing between 0.7 and 1.3 kg. The largest is the male mandrill, around 70 cm in length, and weighing up to 50 kg. Old ...
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Photopsin
Vertebrate visual opsins are a subclass of ciliary opsins and mediate vision in vertebrates. They include the opsins in human rod and cone cells. They are often abbreviated to ''opsin'', as they were the first opsins discovered and are still the most widely studied opsins. Opsins Opsin refers strictly to the apoprotein (without bound retinal). When an opsin binds retinal to form a holoprotein, it is referred to as Retinylidene protein. However, the distinction is often ignored, and opsin may refer loosely to both (regardless of whether retinal is bound). Opsins are G-protein-coupled receptors (GPCRs) and must bind retinal ⁠— typically 11-''cis''-retinal ⁠— in order to be photosensitive, since the retinal acts as the chromophore. When the Retinylidene protein absorbs a photon, the retinal isomerizes and is released by the opsin. The process that follows the isomerization and renewal of retinal is known as the visual cycle. Free 11-''cis''-retinal is photosensitive an ...
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Spectral Sensitivity
Spectral sensitivity is the relative efficiency of detection, of light or other signal, as a function of the frequency or wavelength of the signal. In visual neuroscience, spectral sensitivity is used to describe the different characteristics of the photopigments in the rod cells and cone cells in the retina of the eye. It is known that the rod cells are more suited to scotopic vision and cone cells to photopic vision, and that they differ in their sensitivity to different wavelengths of light. It has been established that the maximum spectral sensitivity of the human eye under daylight conditions is at a wavelength of 555  nm, while at night the peak shifts to 507 nm. In photography, film and sensors are often described in terms of their spectral sensitivity, to supplement their characteristic curves that describe their responsivity. A database of camera spectral sensitivity is created and its space analyzed. For X-ray films, the spectral sensitivity is chosen to b ...
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Photopsin
Vertebrate visual opsins are a subclass of ciliary opsins and mediate vision in vertebrates. They include the opsins in human rod and cone cells. They are often abbreviated to ''opsin'', as they were the first opsins discovered and are still the most widely studied opsins. Opsins Opsin refers strictly to the apoprotein (without bound retinal). When an opsin binds retinal to form a holoprotein, it is referred to as Retinylidene protein. However, the distinction is often ignored, and opsin may refer loosely to both (regardless of whether retinal is bound). Opsins are G-protein-coupled receptors (GPCRs) and must bind retinal ⁠— typically 11-''cis''-retinal ⁠— in order to be photosensitive, since the retinal acts as the chromophore. When the Retinylidene protein absorbs a photon, the retinal isomerizes and is released by the opsin. The process that follows the isomerization and renewal of retinal is known as the visual cycle. Free 11-''cis''-retinal is photosensitive an ...
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Visual Cycle
The visual cycle is a process in the retina that replenishes the molecule retinal for its use in vision. Retinal is the chromophore of most visual opsins, meaning it captures the photons to begin the phototransduction cascade. When the proton is absorbed, the ''11-cis retinal'' photoisomerizes into ''all-trans retinal'' as it is ejected from the opsin protein. Each molecule of retinal must travel from the photoreceptor cell to the RPE and back in order to be refreshed and combined with another opsin. This closed enzymatic pathway of ''11-cis retinal'' is sometimes called Wald's visual cycle after George Wald (1906–1997), who received the Nobel Prize in 1967 for his work towards its discovery. Retinal Retinal is a chromophore that forms photosensitive Retinylidene proteins when covalently bound to proteins called opsins. Retinal can be photoisomerized by itself, but requires to be bound to an opsin protein to both trigger the phototransduction cascade and tune the spectral ...
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Opsin
Animal opsins are G-protein-coupled receptors and a group of proteins made light-sensitive via a chromophore, typically retinal. When bound to retinal, opsins become Retinylidene proteins, but are usually still called opsins regardless. Most prominently, they are found in photoreceptor cells of the retina. Five classical groups of opsins are involved in Visual perception, vision, mediating the conversion of a photon of light into an electrochemical signal, the first step in the Visual phototransduction, visual transduction cascade. Another opsin found in the mammalian retina, melanopsin, is involved in circadian rhythms and Pupillary light reflex, pupillary reflex but not in vision. Humans have in total nine opsins. Beside vision and light perception, opsins may also sense temperature, sound, or chemicals. Structure and function Animal opsins detect light and are the molecules that allow us to see. Opsins are G-protein-coupled receptors (GPCRs), which are chemoreceptors and hav ...
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Photopic Vision
Photopic vision is the vision of the eye under well-lit conditions (luminance levels from 10 to 108  cd/m2). In humans and many other animals, photopic vision allows color perception, mediated by cone cells, and a significantly higher visual acuity and temporal resolution than available with scotopic vision. The human eye uses three types of cones to sense light in three bands of color. The biological pigments of the cones have maximum absorption values at wavelengths of about 420 nm (blue), 534 nm (bluish-green), and 564 nm (yellowish-green). Their sensitivity ranges overlap to provide vision throughout the visible spectrum. The maximum efficacy is 683 lm/W at a wavelength of 555 nm (green). By definition, light at a frequency of hertz has a luminous efficacy of 683 lm/W. The wavelengths for when a person is in photopic vary with the intensity of light. For the blue-green region (500 nm), 50% of the light reaches the image point of the retina. ...
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Rod Cell
Rod cells are photoreceptor cells in the retina of the eye that can function in lower light better than the other type of visual photoreceptor, cone cells. Rods are usually found concentrated at the outer edges of the retina and are used in peripheral vision. On average, there are approximately 92 million rod cells (vs ~6 million cones) in the human retina. Rod cells are more sensitive than cone cells and are almost entirely responsible for night vision. However, rods have little role in color vision, which is the main reason why colors are much less apparent in dim light. Structure Rods are a little longer and leaner than cones but have the same basic structure. Opsin-containing disks lie at the end of the cell adjacent to the retinal pigment epithelium, which in turn is attached to the inside of the eye. The stacked-disc structure of the detector portion of the cell allows for very high efficiency. Rods are much more common than cones, with about 120 million rod cells compar ...
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