Europejara
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Europejara
''Europejara'' is a genus of tapejarid pterosaur from the Early Cretaceous period of Spain. The type species, type and only species known is ''Europejara olcadesorum''. Discovery and naming In 2012, the type species ''Europejara olcadesorum'' was named and described by Romain Vullo, Jesús Marugán-Lobón, Alexander Kellner, Angela Buscalioni, Bernard Gomez, Montserrat de la Fuente and José Moratalla. The generic name combines the names of Europe and the related genus ''Tapejara (pterosaur), Tapejara'', in reference to the fact that ''Europejara'' is the first tapejarid found in that continent. The specific name (zoology), specific name refers to the Olcades, the Celtiberians, Celtiberic tribe inhabiting the region of Province of Cuenca (Spain), Cuenca, the location of the find, in Classical antiquity, Antiquity. The holotype, MCCM-LH 9413, was uncovered at the Las Hoyas site in a chalkstone layer of the Calizas de La Huergina Formation dating from the late Barremian. It cons ...
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Europejara Jaw
''Europejara'' is a genus of tapejarid pterosaur from the Early Cretaceous period of Spain. The type species, type and only species known is ''Europejara olcadesorum''. Discovery and naming In 2012, the type species ''Europejara olcadesorum'' was named and described by Romain Vullo, Jesús Marugán-Lobón, Alexander Kellner, Angela Buscalioni, Bernard Gomez, Montserrat de la Fuente and José Moratalla. The generic name combines the names of Europe and the related genus ''Tapejara (pterosaur), Tapejara'', in reference to the fact that ''Europejara'' is the first tapejarid found in that continent. The specific name (zoology), specific name refers to the Olcades, the Celtiberians, Celtiberic tribe inhabiting the region of Province of Cuenca (Spain), Cuenca, the location of the find, in Classical antiquity, Antiquity. The holotype, MCCM-LH 9413, was uncovered at the Las Hoyas site in a chalkstone layer of the Calizas de La Huergina Formation dating from the late Barremian. It cons ...
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Europejara Skull
''Europejara'' is a genus of tapejarid pterosaur from the Early Cretaceous period of Spain. The type and only species known is ''Europejara olcadesorum''. Discovery and naming In 2012, the type species ''Europejara olcadesorum'' was named and described by Romain Vullo, Jesús Marugán-Lobón, Alexander Kellner, Angela Buscalioni, Bernard Gomez, Montserrat de la Fuente and José Moratalla. The generic name combines the names of Europe and the related genus '' Tapejara'', in reference to the fact that ''Europejara'' is the first tapejarid found in that continent. The specific name refers to the Olcades, the Celtiberic tribe inhabiting the region of Cuenca, the location of the find, in Antiquity. The holotype, MCCM-LH 9413, was uncovered at the Las Hoyas site in a chalkstone layer of the Calizas de La Huergina Formation dating from the late Barremian. It consists of a partial skull with lower jaws, compressed on a slab and counterslab. Two elements of the hyoid are pre ...
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Europejara Olcadesorum
''Europejara'' is a genus of tapejarid pterosaur from the Early Cretaceous period of Spain. The type and only species known is ''Europejara olcadesorum''. Discovery and naming In 2012, the type species ''Europejara olcadesorum'' was named and described by Romain Vullo, Jesús Marugán-Lobón, Alexander Kellner, Angela Buscalioni, Bernard Gomez, Montserrat de la Fuente and José Moratalla. The generic name combines the names of Europe and the related genus '' Tapejara'', in reference to the fact that ''Europejara'' is the first tapejarid found in that continent. The specific name refers to the Olcades, the Celtiberic tribe inhabiting the region of Cuenca, the location of the find, in Antiquity. The holotype, MCCM-LH 9413, was uncovered at the Las Hoyas site in a chalkstone layer of the Calizas de La Huergina Formation dating from the late Barremian. It consists of a partial skull with lower jaws, compressed on a slab and counterslab. Two elements of the hyoid are pre ...
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Tapejarid
Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primitive genera were found in China, indicating that the family has an Asian origin. Description Tapejarids were small to medium-sized pterosaurs with several unique, shared characteristics, mainly relating to the skull. Most tapejarids possessed a bony crest arising from the snout (formed mostly by the premaxillary bones of the upper jaw tip). In some species, this bony crest is known to have supported an even larger crest of softer, fibrous tissue that extends back along the skull. Tapejarids are also characterized by their large nasoantorbital fenestra, the main opening in the skull in front of the eyes, which spans at least half the length of the entire skull in this family. Their eye sockets were small and pear-shaped. Studies of tapejarid ...
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Tapejara (pterosaur)
''Tapejara'' (from a Tupi word meaning "the old being") is a genus of Brazilian pterosaur from the Cretaceous Period (Santana Formation, dating to about 127 to 112 million years ago). ''Tapejara'' crests consisted of a semicircular crest over the snout, and a bony prong which extended back behind the head. It was a small pterosaur, with a wingspan of approximately . Species and classification The type species and only one currently recognized as valid by most researchers, is ''T. wellnhoferi''. The specific name honors German paleontologist Peter Wellnhofer. Two larger species, originally named ''Tapejara imperator'' and ''Tapejara navigans'', were later classified in the genus ''Tapejara''. However, several studies have shown that ''T. imperator'' and ''T. navigans'' are significantly different from ''T. wellnhoferi'' and therefore were reclassified into new genera. The species ''T. imperator'' was given its own genus, ''Tupandactylus'', by Alexander Kellner and Diogenes de A ...
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Azhdarchoid Mandibles
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolutionary ...
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Alexander Kellner
Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, including extinct dinosaurs and crocodylomorphs. Kellner has over 500 publications to his name, has published more than 160 primary studies and two science books. He has participated in paleontological expeditions to many locations including Brazil, Chile, Iran, the United States, Argentina, China, and Antarctica. His scientific achievements include the description of more than thirty species. For his work he has received several honors and prizes, including the TWAS Prize for Earth Sciences from The World Academy of Sciences and admission to the National Order of Scientific Merit (class Comendador), one Brazil's most prestigious awards. Biography Kellner was born in Vaduz, Liechtenstein, son of a German father and Austrian mother. In hi ...
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Romain Vullo
Romain may refer to: People Given name * Romain Bussine (1830–1899), French poet and voice professor * Romain Rolland (1866–1944), French writer * Romain de Tirtoff (1892–1990), French artist and designer known as Erté * Romain Bellenger (1894–1981), French road racing cyclist * Romain Gijssels (1907–1978), Belgian professional road bicycle racer * Romain Maes (1912–1983), Belgian cyclist * Romain Gary (1914–1980), French novelist, film director, World War II pilot, and diplomat * Romain Weingarten (1926–2006), French playwright * Romain Duris (born 1974), French actor * Romain Sardou (born 1974), successful French novelist * Romain Barnier (born 1976), freestyle swimmer * Romain Ferrier (born 1976), French defender * Romain Larrieu (born 1976), goalkeeper * Romain Haguenauer (born 1976), French ice dancing coach, choreographer, and former competitor * Romain Dumas (born 1977), French racing driver * Romain Pitau (born 1977), French football midfielder * Romain ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Mandibula
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone of the skull (discounting the ossicles of the middle ear). It is connected to the temporal bones by the temporomandibular joints. The bone is formed in the fetus from a fusion of the left and right mandibular prominences, and the point where these sides join, the mandibular symphysis, is still visible as a faint ridge in the midline. Like other symphyses in the body, this is a midline articulation where the bones are joined by fibrocartilage, but this articulation fuses together in early childhood.Illustrated Anatomy of the Head and Neck, Fehrenbach and Herring, Elsevier, 2012, p. 59 The word "mandible" derives from the Latin word ''mandibula'', "jawbone" (literally "one used for chewing"), from '' mandere'' "to chew" and ''-bula'' (ins ...
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Autapomorphies
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to the focal taxon (which may be a species, family or in general any clade). It can therefore be considered an apomorphy in relation to a single taxon. The word ''autapomorphy'', first introduced in 1950 by German entomologist Willi Hennig, is derived from the Greek words αὐτός, ''autos'' "self"; ἀπό, ''apo'' "away from"; and μορφή, ''morphḗ'' = "shape". Discussion Because autapomorphies are only present in a single taxon, they do not convey information about relationship. Therefore, autapomorphies are not useful to infer phylogenetic relationships. However, autapomorphy, like synapomorphy and plesiomorphy is a relative concept depending on the taxon in question. An autapomorphy at a given level may well be a synapomorphy at ...
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