Eotitanosuchia
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Eotitanosuchia
Eotitanosuchidae is an extinct family of biarmosuchian therapsids. The Eotitanosuchidae were large predatory therapsids of the Wordian epoch. It was once considered to belong to a separate infraorder of therapsids called Eotitanosuchia. Characteristics The Eotitanosuchians seem to be more advanced than the Biarmosuchia in that the temporal opening behind the eye socket—although small—is still somewhat larger than the biarmosuchians; it is expanded in the upper rear ( posterodorsal) margin, allowing the area of attachment of the adductor ( jaw closing) muscles to be visible from the dorsal (top) view looking down. The eotitanosuchian bite was stronger and more efficient than the biarmosuchian bite. For this reason, some paleontologists see the eotitanosuchids as transitional between the biarmosuchians and higher therapsids. It is at least as likely that features of a larger temporal opening—and hence increased muscle mass and biting power—evolved simultaneously among a nu ...
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Biarmosuchians
Biarmosuchians are an extinct clade of non-mammalian synapsids from the Permian. They are the most basal group of the therapsids. All of them were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation. Characteristics The biarmosuchian skull is very similar to the sphenacodontid skull, differing only in the larger temporal fenestra (although these are still small relative to later therapsids), slightly backward-sloping occiput (the reverse of the pelycosaur condition), reduced number of teeth, and single large canine teeth in both upper and lower jaws, and other features (Carroll 1988 pp. 370, Benton 2000 p. 114). In later specialised Biarmosuchia, these resemble the enlarged canines of the Gorgonopsia. The pre ...
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Biarmosuchia
Biarmosuchians are an extinct clade of non-mammalian synapsids from the Permian. They are the most basal group of the therapsids. All of them were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation. Characteristics The biarmosuchian skull is very similar to the sphenacodontid skull, differing only in the larger temporal fenestra (although these are still small relative to later therapsids), slightly backward-sloping occiput (the reverse of the pelycosaur condition), reduced number of teeth, and single large canine teeth in both upper and lower jaws, and other features (Carroll 1988 pp. 370, Benton 2000 p. 114). In later specialised Biarmosuchia, these resemble the enlarged canines of the Gorgonopsia. The pr ...
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Biarmosuchia
Biarmosuchians are an extinct clade of non-mammalian synapsids from the Permian. They are the most basal group of the therapsids. All of them were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation. Characteristics The biarmosuchian skull is very similar to the sphenacodontid skull, differing only in the larger temporal fenestra (although these are still small relative to later therapsids), slightly backward-sloping occiput (the reverse of the pelycosaur condition), reduced number of teeth, and single large canine teeth in both upper and lower jaws, and other features (Carroll 1988 pp. 370, Benton 2000 p. 114). In later specialised Biarmosuchia, these resemble the enlarged canines of the Gorgonopsia. The pr ...
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Eotitanosuchus
''Eotitanosuchus'' ("dawn giant crocodile") is an extinct genus of biarmosuchian therapsids whose fossils were found in the town of Ochyor in Perm Krai, Russia. It lived about 267 million years ago. The only species is ''Eotitanosuchus olsoni''. Description ''Eotitanosuchus'' is known from a single large skull without a lower jaw. The skull was , but the overall length may have been over , possibly up to and more than in weight for adult specimens. Like ''Biarmosuchus tener'', it was primitive in that, though it was a predator, the temple opening behind the eye was small, giving it a weak bite. The temple was, however, larger at the top than in other biarmosuchians. Paleoecology ''Eotitanosuchus'' fossils were found in the Perm (or Cis-Urals) region of Russia. ''Eotitanosuchus'' was without doubt a dominant animal of its environment. Found preserved in flood deposits (once coastal bogs) containing many skeletons of estemmenosuchids, it has been suggested that this large preda ...
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Eotitanosuchus Olsoni
''Eotitanosuchus'' ("dawn giant crocodile") is an extinct genus of biarmosuchian therapsids whose fossils were found in the town of Ochyor in Perm Krai, Russia. It lived about 267 million years ago. The only species is ''Eotitanosuchus olsoni''. Description ''Eotitanosuchus'' is known from a single large skull without a lower jaw. The skull was , but the overall length may have been over , possibly up to and more than in weight for adult specimens. Like '' Biarmosuchus tener'', it was primitive in that, though it was a predator, the temple opening behind the eye was small, giving it a weak bite. The temple was, however, larger at the top than in other biarmosuchians. Paleoecology ''Eotitanosuchus'' fossils were found in the Perm (or Cis-Urals) region of Russia. ''Eotitanosuchus'' was without doubt a dominant animal of its environment. Found preserved in flood deposits (once coastal bogs) containing many skeletons of estemmenosuchids, it has been suggested that this large pre ...
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Wordian
In the geologic timescale, the Wordian is an age or stage of the Permian. It is the middle of three subdivisions of the Guadalupian Epoch or Series. The Wordian lasted between and million years ago (Ma). It was preceded by the Roadian and followed by the Capitanian. Stratigraphy The Wordian Stage was introduced into scientific literature by Johan August Udden in 1916 and was named after the Word Formation of the North American Permian Basin. The Wordian was first used as a stratigraphic subdivision of the Guadalupian in 1961, when both names were still only used regionally in the southern US. The stage was added to the internationally used ISC timescale in 2001.; 1999: ''Proposal of Guadalupian and Component Roadian, Wordian and Capitanian Stages as International Standards for the Middle Permian Series'', Permophiles 34: pp 3–11. The base of the Wordian Stage is defined as the place in the stratigraphic record where fossils of conodont species ''Jinogondolella aserrata'' fi ...
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Muscles Of Mastication
There are four classical muscles of mastication. During mastication, three muscles of mastication (''musculi masticatorii'') are responsible for adduction of the jaw, and one (the lateral pterygoid) helps to abduct it. All four move the jaw laterally. Other muscles, usually associated with the hyoid, such as the mylohyoid muscle, are responsible for opening the jaw in addition to the lateral pterygoid. Structure The muscles are: * The masseter (composed of the superficial and deep head) * The temporalis (the sphenomandibularis is considered a part of the temporalis by some sources, and a distinct muscle by others) * The medial pterygoid * The lateral pterygoid In humans, the mandible, or lower jaw, is connected to the temporal bone of the skull via the temporomandibular joint. This is an extremely complex joint which permits movement in all planes. The muscles of mastication originate on the skull and insert into the mandible, thereby allowing for jaw movements during contr ...
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Guadalupian Extinctions
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Sphenacodont
Sphenacodontia is a stem-based clade of derived synapsids. It was defined by Amson and Laurin (2011) as "the largest clade that includes ''Haptodus baylei'', ''Haptodus garnettensis'' and '' Sphenacodon ferox'', but not ''Edaphosaurus pogonias''". They first appear during the Late Pennsylvanian (Upper Carboniferous) epoch. From the end of the Carboniferous to the end of the Permian, most of them remained large, with only some secondarily becoming small in size. Basal Sphenacodontia constitute a transitional evolutionary series from early pelycosaurs to ancestral therapsids (which in turn were the ancestors of more advanced forms and finally the mammals). One might say that the sphenacodontians are proto-therapsids (even though there is almost a 30-million-years gap between the separation of the ancestors of therapsids from other sphenacodontians and the first appearance of therapsids in the fossil record). Characteristics The defining characteristics include a thickening of ...
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Cladistic
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Biting
Biting is a common zoological behavior involving the active, rapid closing of the jaw around an object. This behavior is found in toothed animals such as mammals, reptiles, amphibians and fish, but can also exist in arthropods. Myocytic contraction of the muscles of mastication is responsible for generating the force that initiates the preparatory jaw abduction (opening), then rapidly adducts (closes) the jaw and moves the top and bottom teeth towards each other, resulting in the forceful action of a bite. Biting is one of the main functions in most macro-organisms' life, providing them the ability to forage, hunt, eat, build, play, fight and protect, and much more. Biting may be a form of physical aggression due to predatory or territorial intentions, but can also be a normal activity of an animal as it eats, carries objects, softens and prepares food for its young, removes ectoparasites or irritating foreign objects (e.g. burred plant seeds) from body surface, scratches ...
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Muscle Mass
Skeletal muscles (commonly referred to as muscles) are organs of the vertebrate muscular system and typically are attached by tendons to bones of a skeleton. The muscle cells of skeletal muscles are much longer than in the other types of muscle tissue, and are often known as muscle fibers. The muscle tissue of a skeletal muscle is striated – having a striped appearance due to the arrangement of the sarcomeres. Skeletal muscles are voluntary muscles under the control of the somatic nervous system. The other types of muscle are cardiac muscle which is also striated and smooth muscle which is non-striated; both of these types of muscle tissue are classified as involuntary, or, under the control of the autonomic nervous system. A skeletal muscle contains multiple fascicles – bundles of muscle fibers. Each individual fiber, and each muscle is surrounded by a type of connective tissue layer of fascia. Muscle fibers are formed from the fusion of developmental myoblasts in a pr ...
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